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Group 12: Shrubs, trees, or woody vines; perianth in one whorl or absent; flowers bisexual or unisexual; inflorescence various but not catkin or catkin-like spike; pistil 1
1' Plant rooted in soil, not parasitic2. Inflorescence a densespikeor a spheric1. In flowering plants excluding Asteraceae, Cyperaceae, Poaceae, and some other groups, an unbranched inflorescence in which the flowers are sessile and nearly always open from the bottom to the top of the inflorescence. 2. In Asteraceae, Cyperaceae, Poaceae, and some other groups, a spike-like inflorescence is one in which the inflorescence units (e.g., heads in Asteraceae; spikelets in Cyperaceae and Poaceae), instead of individual flowers, are sessile and attached directly to the main axis of the inflorescence, not to branches, and in which floral development may or may not proceed as in 1.
head, or flowers enclosed within a fleshy, hollow receptacle1. A dense, often spheric inflorescence of sessile or subsessile flowers.
2. In Asteraceae and some other groups, a head-like inflorescence is one in which sessile or subsessile inflorescence units (e.g., heads in Asteraceae, umbels enclosed by involucres in Eriogonum), instead of individual flowers, are attached in a short dense cluster without an evident axis or branches.3. Leaves pinnately compound or modified intolineartoElongate, with nearly parallel sides; narrower than elliptic or oblong.
oblongor narrowlyLonger than wide, with nearly parallel sides; wider than linear.
elliptic,In the shape of a flattened circle or ellipse; wider than linear
entire, flattened axes without leaflets; fruit a legume — perianth actually of 2 whorls, the calyx easily overlooked ..... FABACEAE (Mimosoideae)Having margins that are continuous and smooth (i.e., without teeth, lobes, etc.).
3' Leavessimple,Composed of a single part; undivided; unbranched.
ovateor wider in overall outline,Egg-shaped (i.e., widest below the middle) in two dimensions (i.e., in one plane), as a leaf.
entire, toothed, or palmately lobed; fruit not a legumeHaving margins that are continuous and smooth (i.e., without teeth, lobes, etc.).
4. Inflorescence a hollow receptacle enclosing many tiny flowers; ripened inflorescence an externally smooth, fleshy multiple fruit [a fig]; milky latex present ..... MORACEAE (Ficus)4' Inflorescence a short, densespikeor a1. In flowering plants excluding Asteraceae, Cyperaceae, Poaceae, and some other groups, an unbranched inflorescence in which the flowers are sessile and nearly always open from the bottom to the top of the inflorescence. 2. In Asteraceae, Cyperaceae, Poaceae, and some other groups, a spike-like inflorescence is one in which the inflorescence units (e.g., heads in Asteraceae; spikelets in Cyperaceae and Poaceae), instead of individual flowers, are sessile and attached directly to the main axis of the inflorescence, not to branches, and in which floral development may or may not proceed as in 1.
head; ripened pistillate inflorescence an externally roughened spike or spheric head; milky latex present or 01. A dense, often spheric inflorescence of sessile or subsessile flowers.
2. In Asteraceae and some other groups, a head-like inflorescence is one in which sessile or subsessile inflorescence units (e.g., heads in Asteraceae, umbels enclosed by involucres in Eriogonum), instead of individual flowers, are attached in a short dense cluster without an evident axis or branches.5. Perianth parts fused into a tube; flowersbisexual..... ASTERACEAE (2) {G6,7,8,9,14,17}Both male and female reproductive parts occurring and functional in the same plant or structure (e.g., flower, spikelet, inflorescence).
5' Perianth parts free; flowers unisexual6. Lf-bladeentireto toothed, sometimes irregularly few-lobed, faces glabrous or sparsely short-hairy; ripened pistillate inflorescence a fleshyHaving margins that are continuous and smooth (i.e., without teeth, lobes, etc.).
spikeor1. In flowering plants excluding Asteraceae, Cyperaceae, Poaceae, and some other groups, an unbranched inflorescence in which the flowers are sessile and nearly always open from the bottom to the top of the inflorescence. 2. In Asteraceae, Cyperaceae, Poaceae, and some other groups, a spike-like inflorescence is one in which the inflorescence units (e.g., heads in Asteraceae; spikelets in Cyperaceae and Poaceae), instead of individual flowers, are sessile and attached directly to the main axis of the inflorescence, not to branches, and in which floral development may or may not proceed as in 1.
head; milky latex present;1. A dense, often spheric inflorescence of sessile or subsessile flowers.
2. In Asteraceae and some other groups, a head-like inflorescence is one in which sessile or subsessile inflorescence units (e.g., heads in Asteraceae, umbels enclosed by involucres in Eriogonum), instead of individual flowers, are attached in a short dense cluster without an evident axis or branches.petiolebase not expanded, not coveringLeaf stalk, connecting leaf blade to stem; sometimes more or less indistinct.
axillaryPertaining to or within an axil, especially a leaf axil.
bud; stipules membranous, free, generally early-deciduous ..... MORACEAE {G7,11}1. An incompletely developed, more or less embryonic shoot, usually covered with bud scales. 2. An unopened flower, often protected by sepals.
6' Lf-blade ± deeply palmately lobed, thelobes1. A major expansion or bulge, such as on the margin of a leaf, sepal, or petal, or on the surface of an ovary. 2. The free tips of otherwise fused structures, such as sepals or petals; larger than teeth.
entireor ± toothed, faces densely stellate-tomentose when young; ripened pistillate inflorescence a dryHaving margins that are continuous and smooth (i.e., without teeth, lobes, etc.).
headof long-stiff-hairy achenes; milky latex 0;1. A dense, often spheric inflorescence of sessile or subsessile flowers.
2. In Asteraceae and some other groups, a head-like inflorescence is one in which sessile or subsessile inflorescence units (e.g., heads in Asteraceae, umbels enclosed by involucres in Eriogonum), instead of individual flowers, are attached in a short dense cluster without an evident axis or branches.petiolebase expanded, coveringLeaf stalk, connecting leaf blade to stem; sometimes more or less indistinct.
axillaryPertaining to or within an axil, especially a leaf axil.
bud; stipules large and green, fused around stem, persistent or deciduous — pistils actually > 1 ..... PLATANACEAE {G7,9,10}1. An incompletely developed, more or less embryonic shoot, usually covered with bud scales. 2. An unopened flower, often protected by sepals.
2' Inflorescence not a densespikeor a spheric1. In flowering plants excluding Asteraceae, Cyperaceae, Poaceae, and some other groups, an unbranched inflorescence in which the flowers are sessile and nearly always open from the bottom to the top of the inflorescence. 2. In Asteraceae, Cyperaceae, Poaceae, and some other groups, a spike-like inflorescence is one in which the inflorescence units (e.g., heads in Asteraceae; spikelets in Cyperaceae and Poaceae), instead of individual flowers, are sessile and attached directly to the main axis of the inflorescence, not to branches, and in which floral development may or may not proceed as in 1.
head, flowers not enclosed in a fleshy, hollow receptacle1. A dense, often spheric inflorescence of sessile or subsessile flowers.
2. In Asteraceae and some other groups, a head-like inflorescence is one in which sessile or subsessile inflorescence units (e.g., heads in Asteraceae, umbels enclosed by involucres in Eriogonum), instead of individual flowers, are attached in a short dense cluster without an evident axis or branches.7. Flowers unisexual, plant monoecious; pistillate inflorescence 1–5-flowered, maturing as a bur with spiny, knob-like, or membranousbracttips; staminate inflorescence aReduced, leaf- or scale-like structure subtending a branch, cone scale, peduncle, pedicel, or flower.
head..... ASTERACEAE (Ambrosia)1. A dense, often spheric inflorescence of sessile or subsessile flowers.
2. In Asteraceae and some other groups, a head-like inflorescence is one in which sessile or subsessile inflorescence units (e.g., heads in Asteraceae, umbels enclosed by involucres in Eriogonum), instead of individual flowers, are attached in a short dense cluster without an evident axis or branches.7' Flowersbisexual, or if unisexual, pistillate inflorescence not 1–5-flowered and maturing as a bur with spiny, knob-like, or membranousBoth male and female reproductive parts occurring and functional in the same plant or structure (e.g., flower, spikelet, inflorescence).
bracttips; staminate inflorescence variousReduced, leaf- or scale-like structure subtending a branch, cone scale, peduncle, pedicel, or flower.
8. Ovary inferior, partly so, or appearing so9. Herbage covered with silvery scales; fruit drupe-like — ovary actually superior, ripening as anachene, tightly enclosed byDry, indehiscent, 1-seeded fruit from a 1-chambered ovary, sometimes winged, often appearing to be a naked seed. A 1-seeded dry fruit derived from an inferior ovary of > 1 carpel (e.g., Asteraceae, Dipsacaceae) is sometimes called a cypsela.
hypanthiumbase that becomes fleshy at maturity ..... ELAEAGNACEAEStructure generally in the shape of a tube, cup, or bowl, derived from the fused lower portions of the perianth and stamens, from which these parts seem to arise, and to which the ovary wall is fused in an inferior ovary (to which the ovary wall is partially fused in a half-inferior ovary; from which the ovary is free in a superior ovary).
9' Herbage without silvery scales; fruit various10. Inflorescence unit an involucredhead, superficially resembling a single flower ..... ASTERACEAE (2) {G6,7,8,9,14,17}1. A dense, often spheric inflorescence of sessile or subsessile flowers.
2. In Asteraceae and some other groups, a head-like inflorescence is one in which sessile or subsessile inflorescence units (e.g., heads in Asteraceae, umbels enclosed by involucres in Eriogonum), instead of individual flowers, are attached in a short dense cluster without an evident axis or branches.10' Inflorescence open or flowers solitary or paired11. Trees, evergreen; stamens many; leaves aromatic — perianth early-deciduous as cap ..... MYRTACEAE (Eucalyptus)11' Shrubs or woody vines, evergreen or deciduous; stamens 4–12; leaves generally not aromatic12. Perianthrotate,Wheel-shaped, spreading, or saucer-shaped; often applied to a fused corolla with a short or nonexistent tube and a spreading limb.
lobes>> tube; leaves1. A major expansion or bulge, such as on the margin of a leaf, sepal, or petal, or on the surface of an ovary. 2. The free tips of otherwise fused structures, such as sepals or petals; larger than teeth.
whorled; fruit 2-lobed ..... RUBIACEAE (Galium) {G6,7,8,9}Arranged in groups of three or more at nodes or positions along an axis (e.g., three leaves per node).
12' Perianth tubular,lobes<< tube; leaves1. A major expansion or bulge, such as on the margin of a leaf, sepal, or petal, or on the surface of an ovary. 2. The free tips of otherwise fused structures, such as sepals or petals; larger than teeth.
alternateor1. Arranged singly, often spirally, along an axis - e.g., one leaf per node. 2. Occurring in different ranks, appearing to be between, not directly above or below, as 'stamens alternate petals'.
opposite; fruit not lobed1. Arranged in pairs along an axis - e.g., two leaves per node. 2. Occurring in the same rank, directly above or below, as 'stamens opposite petals'. 3. Located directly across from.
13. Leavesalternate; perianth tube U-shaped; twining vine ..... ARISTOLOCHIACEAE (Aristolochia)1. Arranged singly, often spirally, along an axis - e.g., one leaf per node. 2. Occurring in different ranks, appearing to be between, not directly above or below, as 'stamens alternate petals'.
13' Leavesopposite; perianth tube straight;1. Arranged in pairs along an axis - e.g., two leaves per node. 2. Occurring in the same rank, directly above or below, as 'stamens opposite petals'. 3. Located directly across from.
erectshrub ..... CAPRIFOLIACEAE (Lonicera involucrata)Upright; vertically oriented.
8' Ovary superior14. Leavesoppositeor1. Arranged in pairs along an axis - e.g., two leaves per node. 2. Occurring in the same rank, directly above or below, as 'stamens opposite petals'. 3. Located directly across from.
whorledArranged in groups of three or more at nodes or positions along an axis (e.g., three leaves per node).
15. Fruit winged,indehiscent; leaves generally lobed or compoundNot opening inherently to release contents; usually pertaining to fruits.
16. Style evident, stigma 1,entireor slightly 2-lobed; fruit 1-winged, not splitting; leaves pinnately compound (leaflet generally 1 in Fraxinus anomala) ..... OLEACEAE (Fraxinus) {G7,9,16}Having margins that are continuous and smooth (i.e., without teeth, lobes, etc.).
16' Style very short, stigmas 2(3), elongate; fruit 2(3)-winged, generally splitting into mericarps; leaves palmately lobed or compound or pinnately compound ..... SAPINDACEAE (Acer) {G7,23}15' Fruit not winged; leavessimple,Composed of a single part; undivided; unbranched.
entireor toothedHaving margins that are continuous and smooth (i.e., without teeth, lobes, etc.).
17. Flowersbisexual; stamens 20–25(30) ..... ROSACEAE (Coleogyne)Both male and female reproductive parts occurring and functional in the same plant or structure (e.g., flower, spikelet, inflorescence).
17' Flowers unisexual; stamens generally 3–12
18' Leaf glabrous or ± hairy
19' Leaves not needle-like, not grooved20. Staminate and pistillate flowers insessileumbels; style 1, stigmaWithout a petiole, peduncle, pedicel, or other kind of stalk.
entireor 2-lobed ..... OLEACEAE (Forestiera) {G7,9}Having margins that are continuous and smooth (i.e., without teeth, lobes, etc.).
20' Staminate flowers clustered, pistillate flowers generally solitary; styles or stigmas (2)3–521. Staminate inflorescence aracemeorIn flowering plants excluding Asteraceae, Cyperaceae, Poaceae, and some other groups, an unbranched inflorescence in which the flowers are borne on pedicels and nearly always open from the bottom to the top of the inflorescence. 2. In Asteraceae, Cyperaceae, Poaceae, and some other groups, a raceme-like inflorescence is one in which the inflorescence units (e.g., heads in Asteraceae; spikelets in Cyperaceae and Poaceae), instead of individual flowers, are stalked and attached directly to the main axis of the inflorescence, not to branches, and in which floral development may or may not proceed as in 1.
panicle, flowers clearly stalked; fruit 3–4-lobed ..... PICRODENDRACEAE {G7,9}1. In flowering plants excluding Asteraceae, Cyperaceae, Poaceae, and some other groups, a branched inflorescence in which the basal or lateral flowers (or some of them) open before the terminal or central flowers on any axis. 2. In Asteraceae, Cyperaceae, Poaceae, and some other groups, a panicle-like inflorescence is one in which at least some of the inflorescence units (e.g., heads in Asteraceae; spikelets in Cyperaceae and Poaceae), instead of individual flowers, are attached (stalked or unstalked) to branches and not directly to the main axis of the inflorescence and in which floral development may or may not proceed as in 1.
21' Staminate inflorescence head-like, flowers subsessile; fruit unlobed ..... SIMMONDSIACEAE {G7,9}14' Leavesalternate1. Arranged singly, often spirally, along an axis - e.g., one leaf per node. 2. Occurring in different ranks, appearing to be between, not directly above or below, as 'stamens alternate petals'.
22. Leaves pinnately compound or very deeply pinnately divided23. Perianth parts ± brown-green, inconspicuous, bract-like; flowers unisexual, plants dioecious ..... ANACARDIACEAE (Pistacia) {G7,11}23' Perianth parts petal-like, conspicuous; flowersbisexualBoth male and female reproductive parts occurring and functional in the same plant or structure (e.g., flower, spikelet, inflorescence).
24. Shrub; flowers solitary; perianthradial, parts 6, white — sepals actually present inDivisible into mirror-image halves in three or more ways.
bud, but early-deciduous, 0 in open flower ..... PAPAVERACEAE (Romneya) {G22}1. An incompletely developed, more or less embryonic shoot, usually covered with bud scales. 2. An unopened flower, often protected by sepals.
24' Tree; flowers inraceme; perianthIn flowering plants excluding Asteraceae, Cyperaceae, Poaceae, and some other groups, an unbranched inflorescence in which the flowers are borne on pedicels and nearly always open from the bottom to the top of the inflorescence. 2. In Asteraceae, Cyperaceae, Poaceae, and some other groups, a raceme-like inflorescence is one in which the inflorescence units (e.g., heads in Asteraceae; spikelets in Cyperaceae and Poaceae), instead of individual flowers, are stalked and attached directly to the main axis of the inflorescence, not to branches, and in which floral development may or may not proceed as in 1.
bilateral, parts 4, yellow or orange, proximally ± red ..... PROTEACEAEDivisible into mirror-image halves in only one way.
25. Inflorescence flower-like with a cup-likeinvolucrebearing colored, often petal-like nectaries; each stamen [actually a staminate flower] with a jointed stalk [composed of a filament and a pedicel]; pistil [actually pistillate flower] 3-lobed, generallyGroup of bracts more or less held together as a unit, subtending a flower, fruit (acorn cup), or inflorescence.
exsertedfrom involucre; milky latex present ..... EUPHORBIACEAE (Euphorbia)Protruding out of surrounding structure(s) (e.g., stamens exserted from corolla).
25' Inflorescences various, but not flower-like; stamens generally > 1 per flower, the filaments not jointed; pistils various; milky latex generally 026. Anthers opening by pores with hinged lids;leafstrongly aromatic — 2-whorled nature of perianth may not be evident in open flowers ..... LAURACEAE {G16}Organ arising from a stem, generally composed of a stalk (petiole) and a flat, expanded, green, photosynthetic area (blade); distinguished from a leaflet by the presence in its axil of a bud, branch, thorn, or flower; sometimes with lateral, basal appendages (stipules); either simple (toothed, lobed, or dissected but not divided into leaflets) or compound (divided into leaflets).
26' Anthers opening by slits;leafgenerally not aromaticOrgan arising from a stem, generally composed of a stalk (petiole) and a flat, expanded, green, photosynthetic area (blade); distinguished from a leaflet by the presence in its axil of a bud, branch, thorn, or flower; sometimes with lateral, basal appendages (stipules); either simple (toothed, lobed, or dissected but not divided into leaflets) or compound (divided into leaflets).
27. Leaves fleshy,cylindricto ± flat ..... CHENOPODIACEAE (Suaeda)Elongate, with parallel sides and, at any point, round in transverse section.
27' Leaves not fleshy, generally flat28. Leaf palmately veined and lobed29. Woody vine; tendrils present — calyx 0 or reduced to anentireor slightly lobed rim, easily overlooked ..... VITACEAE (Vitis) {G7,9,23}Having margins that are continuous and smooth (i.e., without teeth, lobes, etc.).
29' Erect orspreadingshrub; tendrils 0Oriented more or less perpendicularly to the axis of attachment; often, more or less horizontal.
30. Leafpeltate, glabrous; flowers unisexual, perianth parts ± green, free; stamens very many, the filaments irregularly joined in clusters ..... EUPHORBIACEAE (Ricinus) {G7,9}With the stalk attached toward the middle, not at a margin, of a flat structure such as an indusium, scale, or leaf.
30' Leaf notpeltate, stellate-hairy; flowersWith the stalk attached toward the middle, not at a margin, of a flat structure such as an indusium, scale, or leaf.
bisexual, perianth parts yellow or orange to ± red, petal-like; stamens 5, filaments fused in 1 group ..... MALVACEAE (Fremontodendron) {G19}Both male and female reproductive parts occurring and functional in the same plant or structure (e.g., flower, spikelet, inflorescence).
28' Leaf pinnately veined or only midvein evident, generally unlobed31. Perianth parts 2–10 cm; ovules many — sepals actually present inbud, but early-deciduous, 0 in open flower ..... PAPAVERACEAE (Papaveroideae) {G15,20,22,24}1. An incompletely developed, more or less embryonic shoot, usually covered with bud scales. 2. An unopened flower, often protected by sepals.
31' Perianth parts generally < 1.5 cm; ovules 1–2 per chamber32. Plant covered with ± gray or silvery scales33. Flowers tubular in proximal half, some or allbisexual; tree or large shrub; leaves linear-lanceolate toBoth male and female reproductive parts occurring and functional in the same plant or structure (e.g., flower, spikelet, inflorescence).
oblong..... ELAEAGNACEAE (Elaeagnus)Longer than wide, with nearly parallel sides; wider than linear.
33' Flowers not tubular, all unisexual, plant dioecious; shrub; leaves various34. Ovary of pistillate flower enclosed by 2appressedbracts;Parallel or nearly parallel to and often in contact with surface of origin; used to describe the disposition of hairs, leaves, pedicels, etc.
leafsurface scales powder-like, notOrgan arising from a stem, generally composed of a stalk (petiole) and a flat, expanded, green, photosynthetic area (blade); distinguished from a leaflet by the presence in its axil of a bud, branch, thorn, or flower; sometimes with lateral, basal appendages (stipules); either simple (toothed, lobed, or dissected but not divided into leaflets) or compound (divided into leaflets).
stellate..... CHENOPODIACEAE (Atriplex) {G6,8}Pertaining to a hair or other structure with three or more branches radiating in two or three dimensions from a common point.
34' Ovary of pistillate flower subtended by 5 sepals;leafsurface scalesOrgan arising from a stem, generally composed of a stalk (petiole) and a flat, expanded, green, photosynthetic area (blade); distinguished from a leaflet by the presence in its axil of a bud, branch, thorn, or flower; sometimes with lateral, basal appendages (stipules); either simple (toothed, lobed, or dissected but not divided into leaflets) or compound (divided into leaflets).
stellate..... EUPHORBIACEAE (Croton) {G6,8}Pertaining to a hair or other structure with three or more branches radiating in two or three dimensions from a common point.
32' Plant without scales35. Perianth parts 6 — 2-whorled nature of perianth may not be evident in open flowers36. Flowers solitary,axillary, unisexual; fruit a berry ..... ERICACEAE (Empetrum) (2) {G7,9,16}Pertaining to or within an axil, especially a leaf axil.
36' Flowers in involucred umbels,bisexual; fruit anBoth male and female reproductive parts occurring and functional in the same plant or structure (e.g., flower, spikelet, inflorescence).
achene..... POLYGONACEAE (Eriogonum) {G6,8}Dry, indehiscent, 1-seeded fruit from a 1-chambered ovary, sometimes winged, often appearing to be a naked seed. A 1-seeded dry fruit derived from an inferior ovary of > 1 carpel (e.g., Asteraceae, Dipsacaceae) is sometimes called a cypsela.
35' Perianth parts generally 4 or 537. Stamens = in number to andalternatesepals ..... RHAMNACEAE {G21,23}1. Arranged singly, often spirally, along an axis - e.g., one leaf per node. 2. Occurring in different ranks, appearing to be between, not directly above or below, as 'stamens alternate petals'.
37' Stamens = in number to andoppositesepals or > in number than sepals1. Arranged in pairs along an axis - e.g., two leaves per node. 2. Occurring in the same rank, directly above or below, as 'stamens opposite petals'. 3. Located directly across from.
38. Style and stigma 139. Style feathery with long, stiff hairs; fruit anachenesurrounded by a tubularDry, indehiscent, 1-seeded fruit from a 1-chambered ovary, sometimes winged, often appearing to be a naked seed. A 1-seeded dry fruit derived from an inferior ovary of > 1 carpel (e.g., Asteraceae, Dipsacaceae) is sometimes called a cypsela.
hypanthium; branches stiff ..... ROSACEAE (Cercocarpus)Structure generally in the shape of a tube, cup, or bowl, derived from the fused lower portions of the perianth and stamens, from which these parts seem to arise, and to which the ovary wall is fused in an inferior ovary (to which the ovary wall is partially fused in a half-inferior ovary; from which the ovary is free in a superior ovary).
39' Style glabrous; fruit an exposed drupe; branches pliable — corolla 0 or reduced to minute scales, easily overlooked ..... THYMELAEACEAE38' Styles or stigmas 2–440. Fruit a drupe; flowers staminate andbisexual..... CANNABACEAE (Celtis) {G7}Both male and female reproductive parts occurring and functional in the same plant or structure (e.g., flower, spikelet, inflorescence).
40' Fruit a winged nutlet or splitting into 1–2-seeded segments that release seeds; flowers unisexual orbisexualBoth male and female reproductive parts occurring and functional in the same plant or structure (e.g., flower, spikelet, inflorescence).
41' Shrub; fruit splitting into segments42. Leaveselliptic,In the shape of a flattened circle or ellipse; wider than linear
crenate; fruit segments 1-seeded ..... EUPHORBIACEAE (Bernardia)Pertaining to margins with shallow, rounded teeth, between which are usually acute sinuses (i.e., scalloped)
42' Leaves oblanceolate to obovate,entire; fruit segments 1–2-seeded ..... PICRODENDRACEAE (Tetracoccus hallii)Having margins that are continuous and smooth (i.e., without teeth, lobes, etc.).