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Michael H. Nee

Annual to shrub. Leaf: generally simple, generally alternate, generally petioled; stipules 0; blade entire to deeply lobed. Inflorescence: various. Flower: bisexual; calyx lobes generally 5; corolla ± radial, cylindric to rotate, lobes generally 5; stamens 5, on corolla tube, alternate lobes; ovary superior, generally 2-chambered, style 1. Fruit: berry, loculicidal or septicidal capsule, [(drupe)], 2–5-chambered.
75 genera, 3000 species: worldwide, especially ± tropics; many alien weeds in California; many cultivated for food, drugs, or ornamental (potato, tomato, peppers, tobacco, petunia); many TOXIC. [Hunziker 2001 Genera Solanacearum. Koeltz Scientific Books] Nicandra physalodes (L.) Gaertn. is a waif. —Scientific Editor: Thomas J. Rosatti.
Unabridged references: [Hunziker 2001 Genera Solanacearum: The Genera of Solanaceae Illustrated, Arranged According to a New System by A.T. Hunziker. Koeltz Scientific Books, Königstein, Germany]

Key to Solanaceae

Annual to small tree. Leaf: ± basal or not; margin entire, ± wavy or not. Inflorescence: raceme or panicle, terminal; lower bracts leaf-like. Flower: calyx 5-lobed, ± enlarging, not fully enclosing fruit; corolla generally radial, generally funnel-shaped to salverform; stamens 5, equal or 1 smaller. Fruit: septicidal capsule. Seed: many, minute, angled.
± 60 species: generally America; Nicotiana tabacum L. widely cultivated, tobacco of commerce. (J. Nicot, said to have introduced tobacco to Eur, 1530–1600) Seriously TOXIC to livestock.
Unabridged references: [Goodspeed 1954 Chron Bot 16:1–536]

Key to Nicotiana

N. quadrivalvis Pursh
Annual 3–20 dm, glandular-hairy. Leaf: 4–15 cm; basal, lower cauline short-petioled, elliptic to ovate; upper sessile, reduced. Inflorescence: bracts < 35 mm, ± linear. Flower: calyx 15–20 mm, 10-ridged, lobes narrowly lanceolate, 1 > tube or not; corolla ± salverform, white, green- or violet-tinged, tube 25–50 mm, limb 20–50 mm wide; stamens unequal, attached at various levels above tube middle. Fruit: 15–20 mm.
Open, well-drained washes, slopes; < 1500 m. California Floristic Province, Desert; to Washington, central United States. [Nicotiana bigelovii (Torr.) S. Watson; Nicotiana bigelovii var. wallacei A. Gray; Nicotiana quadrivalvis var. bigelovii (Torr.) DeWolf; Nicotiana quadrivalvis var. quadrivalvis] Widely cultivated by western America native people; cultivated plants sometimes recognized, with Nicotiana quadrivalvis var. bigelovii (Torr.) DeWolf for wild plants. May–Oct [Online Interchange]

Previous taxon: Nicotiana obtusifolia
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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora,, accessed on Nov 25 2015
Citation for this treatment: [Author of taxon treatment] 2013. Nicotiana, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on Nov 25 2015

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Geographic subdivisions indicated for the distribution of Nicotiana quadrivalvis Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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Data provided by the participants of the Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.