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HYDRANGEACEAE HYDRANGEA FAMILY

Robert E. Preston & Charles F. Quibell

Perennial herb to small tree or vine. Stem: < 3 m, generally erect; bark generally peeling as thin sheets or narrow strips. Leaf: generally simple, opposite, deciduous or not, ± hairy; stipules 0; blade ± round to narrowly elliptic, entire or toothed. Inflorescence: cyme, raceme, panicle, or flower 1, terminal or axillary, generally bracted. Flower: bisexual, radial, flowers on inflorescence margins occasionally sterile and enlarged; sepals 4–10, free or fused at base, spreading or erect; petals 4–7, free, ± round to narrowly elliptic; stamens 8–12 in 2 whorls or many and clustered, filament base linear or wide and flat; pistil 1, ovary superior to inferior, chambers 2–8, ovules 1–2 or many per chamber, placentas axile or parietal, styles 1–8, free or fused at base. Fruit: capsule, loculicidal or septicidal; styles persistent or not. Seed: generally many, small to minute, oblong to fusiform, winged or not.
18 genera, ± 250 species: generally temperate, subtropical northern hemisphere; some cultivated for ornamental (Carpenteria, Hydrangea, Philadelphus). [Hufford et al. 2001 Int J Plant Sci 162:835–846] Philadelphaceae in TJM (1993). —Scientific Editors: Douglas H. Goldman, Bruce G. Baldwin.
Unabridged references: [Hufford 1997 Int J Plant Sci 158:652–672; Soltis et al. 1995 Amer J Bot 82:504–514]

Key to Hydrangeaceae

PHILADELPHUS MOCK ORANGE
Shrub < 3 m. Stem: bark red-brown, aging gray, peeling as narrow rectangles or strips; twigs glabrous to hairy. Leaf: deciduous, petioled; blade 3-veined from base, ± glabrous to hairy, margin entire to toothed. Inflorescence: flower 1, or raceme to panicle, terminal, ± open. Flower: fragrant; sepals 4–5, glabrous to hairy; petals 4–5, white; stamens generally many, clustered, filaments linear, fused at base; ovary 1/2 to completely inferior, chambers 4–5, placentas axile, ovules many, style 1, branches generally 4, stigmas linear along style branches. Fruit: becoming woody, generally loculicidal. Seed: many, generally fusiform, generally brown.
± 65 species: temperate America, Eurasia. (Greek: for Ptolemy Philadelphus, Greek king of Egypt, 309–247 BC) [Frazier 1999 New Mexico Botanist 13:1–6] Many intergrading infraspecific taxa described; more study needed to determine which warrant recognition.
Unabridged references: [Hitchcock 1943 Madroño 7:35–56; Hu 1956 J Arnold Arbor 37:15–90]

Key to Philadelphus

P. lewisii Pursh WILD MOCK ORANGE
NATIVE
Leaf: petiole 3–8 mm; blade 20–40 mm wide, ovate, margin entire to toothed, flat. Flower: sepals 4–7 mm; petals 8–12 mm, obovate to oblong.
2n=26. Slopes, canyons in conifer and hardwood forest, montane chaparral; 30–2440 m. Northwestern California, Cascade Range, n&c Sierra Nevada, s High Sierra Nevada, Sacramento Valley (Sutter Buttes), San Gabriel Mountains, Peninsular Ranges (Santa Ana Mtns); to British Columbia, Montana. May–Jul [Online Interchange]
Unabridged synonyms: [Philadelphus lewisii subsp. californicus (Benth.) Munz; Philadelphus lewisii subsp. gordonianus (Lindl.) Munz.]

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Next taxon: Philadelphus microphyllus

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Citation for the whole project: Jepson Flora Project (eds.) 2013. Jepson eFlora, http://ucjeps.berkeley.edu/IJM.html, accessed on Nov 22 2014
Citation for this treatment: [Author of taxon treatment] 2013. Philadelphus, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=37691, accessed on Nov 22 2014

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click for enlargement Philadelphus lewisii
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Gladys Lucille Smith © 1999 California Academy of Sciences

Geographic subdivisions indicated for the distribution of Philadelphus lewisii Markers link to CCH specimen records. If the markers are obscured, reload the page [or change window size and reload]. Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).

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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa.
Blue line denotes eFlora flowering time.