|Jepson Flora Project: Jepson Interchange|
|TREATMENT FROM THE JEPSON MANUAL (1993)||
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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California
AND IS MAINTAINED FOR ARCHIVAL PURPOSES ONLY
Annual, perennial herb, sometimes aquatic
Leaves generally basal and cauline, generally alternate, simple or compound; petioles at base generally flat, sometimes sheathing or stipule-like
Inflorescence: cyme, raceme, panicle, or flowers solitary
Flower generally bisexual, radial; sepals generally 5, free, early deciduous or withering in fruit, generally green; petals 0many, free; stamens generally 10many; pistils 1many, ovary superior, chamber 1, style 1, generally ± persistent in fruit as beak, ovules 1many
Fruit: achene, follicle, berry, or utricle-like, 1many-seeded
Genera in family: ± 60 genera, 1700 species: worldwide, especially n temp, tropical mtns; many ornamental (Adonis, Aquilegia, Clematis, Consolida, Delphinium, Erianthis, Helleborus ),
some highly TOXIC (Aconitum, Actaea, Delphinium, Ranunculus )
Reference: [Duncan & Keener 1991 Phytologia 70:2427]
Perennial; root generally < 10 cm, ± fibrous or fleshy; buds generally obscure
Stem generally 1, erect, generally unbranched; base generally ± as wide as root, generally firmly attached to root, generally ± reddish or purplish
Leaves simple, basal and cauline, petioled; blades generally palmately lobed, deep lobes generally 35, generally < 6 mm wide, generally also lobed; lower leaves generally dry, often 0 in flower; cauline merging into bracts upward
Inflorescence: raceme or somewhat branched, terminal; flowers generally 1025; pedicels generally ± spreading
Flower bilateral; sepals 5, petal-like, generally spreading, generally ± dark blue, uppermost spurred; petals 4, << sepals, upper 2 with nectar-secreting spurs enclosed in uppermost sepal, lower 2 clawed, with blades generally 48 mm, notched, generally ± perpendicular to claws, generally colored like sepals, generally obviously hairy; pistils 3(5)
Fruit aggregate of 3(5) erect follicles, generally 2.54 X longer than wide
Seed dark brown to black, often appearing white, generally winged when immature, generally without inflated collar; coat cell margins generally straight
Etymology: (Latin: dolphin, from bud shape)
Reference: [Lewis & Epling 1954 Brittonia 8:122]
Hybrids common, especially in disturbed places. Root length here includes coarse but not thread-like parts. Most species highly TOXIC, attractive and causing many deaths to cattle, less often to horses, sheep.
Horticultural information: Exc as noted, successful In cultivation only within natural range and habitat. Lowland subsp.: DRY. Upland species: winter chilling required.
Root ± spheric, clustered or not
Stem 5100 (generally < 50) cm; base narrower than root, not firmly attached to root, glabrous to puberulent
Leaves mostly on lower 20% of stem, ± glabrous; lobes 725 (6 or more extending > 60% to petiole), < 7 mm wide
Inflorescence: flowers rarely > 12; pedicels 775 mm, 1550 mm apart, glabrous or puberulent
Flower: sepals generally ± reflexed, generally ± glabrous, lateral 818 mm, spur 820 mm
Fruit 717 mm, ± curved
Seed with inflated collar at widest end, otherwise smooth, shiny
Ecology: Open woodlands, sagebrush scrub, meadow edges, streamsides
Elevation: 3003300 m.
Bioregional distribution: Klamath Ranges, Cascade Range, High Sierra Nevada, Modoc Plateau
Distribution outside California: to Oregon, Idaho, Nevada
Synonyms: D. sonnei Greene
Hybridizes with D. depauperatum , D. nudicaule , D. polycladon ; extremely difficult, variable complex.
Horticultural information: DRN, DRY; DFCLT.