CENTER FOR PHYCOLOGICAL DOCUMENTATION
DECEW'S GUIDE

Costaria costata ( C. Agardh ) Saunders
	CONTENTS     FIGURES     ABOUT     INDEXES: Reds · Greens · Browns     KEYS: Browns

habit ( young sporophyte ) Synonyms: Costaria mertensii J. Agardh, C. quadrinervia Ruprecht, C. turneri Greville ( see Taxonomic notes: p. 000 ) Illustrations: Sporophyte: Abbott and Hollenberg 1976, p. 238, fig. 195 ( habit ); Angst 1927 b, p. 305, 307, figs. 24, 25, 27 & ndash; 31 ( development from zygote ); Cameron 1912, pl. XVI, fig. 1 ( photo: habit ); Chihara 1975, p. 73 ( color photo: habit ); Kanda 1936, p. 241, fig. 15 C, D ( development from zygote ); Okada 1956, pl. 48 ( color photo: habit ); Okamura 1928 b, pl. CCXXVI ( habit ); Postels and Ruprecht 1840, pl. XXIV ( habit ); Scagel 1967, p. 131, fig. 51 ( habit ); Segawa 1956, pl. 23, fig. 189 ( color photo: habit ); Setchell and Gardner 1925, pl. 56, fig. A ( photo: habit ), pl. 79, fig. B ( photo: habit ); Smith 1944, pl. 22, fig. 1 ( habit ). Gametophytes: Kanda 1936, p. 238, fig. 12, p. 239, fig. 13 ( development from zoospores ), p. 240, fig. 14 ( male ), p. 241, fig. 15 A, B ( female ); Angst 1927 b, p. 301, figs. 1 & ndash; 11 ( development from zoospores ), p. 303, 305, figs. 12, 19 & ndash; 22 ( male ), figs. 13 & ndash; 17, 23 ( female ) Pacific Coast Distribution: Unalaska I ., Aleutian Is ., Alaska ( Setchell and Gardner 1903, p. 265 ) to Partington Pt ., 16 km south of Pt. Sur, Monterey Co ., Calif. ( Silva 2289 in UC ); Channel Is ., Calif. ( San Miguel I.: Miller in UC ); ( unsubstantiated report for San Pedro, Los Angeles Co ., Calif.: Setchell and Gardner 1925, p. 610 ) Taxonomy: Doty 1947, p. 41; Nagai 1940, p. 92; Setchell and Gardner 1925, p. 610; D. Saunders 1895 Molecular Taxonomy: Saunders and Druehl 1993 b; Druehl and Saunders 1992; Bhattacharya and Druehl 1989; Bhattacharya and Druehl 1988 Evolution: Druehl and Saunders 1992; Bhattacharya et al. 1991; Bhattacharya 1988; Estes and Steinberg 1988 Life History: alternation of heteromorphic phases, with a macroscopic sporophyte and dioecious microscopic gametophytes ( Kanda 1936; Angst 1927 b ) Phenology: Maxell and Miller 1996 Chromosomes: Yasui 1992; Cole 1967 b, p. 523 Genetics: Bhattacharya, Mayes et al. 1991; Bhattacharya 1988; Bhattacharya and Druehl 1987 Biology/Ecology: Hurd and Stevens 1997; Shaffer and Parks 1994; Bhattacharya et al. 1990; L & uuml; ning 1990; L & uuml; ning and Freshwater 1988, p. 312; Druehl and Breen 1986, p. 99; Dayton 1985, p. 220; Foster and Schiel 1985, p. 28; Duggins 1983, p. 1615; Whyte and Englar 1983, p. 163; Turner 1983 a, p. 63; Druehl and Green 1982, p. 167; L & uuml; ning 1982, p. 54; Duggins 1981; Pace 1981, p. 460; Hodgson and Waaland 1980, p. 110; Calvin and Ellis 1978; Druehl and Hsiao 1977; Vadas 1977; Dayton 1975, p. 139; Scagel 1973, p. 132; Hasegawa and Sanbonsuga 1972; Druehl 1970, p. 239; Druehl 1969, p. 163; Druehl and Hsiao 1969; Paine and Vadas 1969 a, p. 714; Paine and Vadas 1969 b, p. 80; Sanbonsuga and Hasegawa 1969; Neushul 1967, p. 85; Sanbonsuga and Hasegawa 1967; R. Lee 1965 a, p. 14; R. Lee 1965 b, p. 111; Widdowson 1965 b, p. 1428; Scagel 1961, p. 528; Stephenson and Stephenson 1961 b, p. 228; Rigg and Miller 1949 Epiphytes: Ceramium gardneri , Hecatonema primarium , Hecatonema streblonematoides , Myrionema corunnae Ultrastructure: Henry and Cole 1982 a, p. 551; Henry and Cole 1982 b, p. 571; Henry 1980, p. 9 Chemistry: Van Alstyne et al. 1999; Nagahisa et al. 1992; Ragan and Glombitza 1986, p. 188; Tsuchiya 1979; Saenko et al. 1978 Physiology: K. Schmitz 1982, p. 171; Kremer 1981, p. 504; Kremer 1979, p. 245; Jolliffe and Tregunna 1970, p. 296 Economic Applications: Druehl and Boal 1981, p. 578; Mumford 1980, p. 19

							J	F	M	A	M	J	J	A	S	O	N	D
BRITISH COLUMBIA						Pl													Common, in pools and on rocks and wood in the low and mid intertidal and upper subtidal in fully sheltered to fully exposed habitats. Scagel 1957, p. 100. Scagel 1973, p. 132. Hawkes et al. 1979, p. 92.
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COLUMBIA RIVER						Pl													Common, throughout the range, in pools and on rocks in the low intertidal and upper subtidal, in protected localities. Doty 1947, p. 41. Sanborn and Doty 1947, p. 30. Markham and Celestino 1977, p. 258.
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CAPE ARAGO						Pl													Uncommon, in pools and on rocks in the low intertidal. Apparently disjunct between Crescent City and Punta Gorda. ( Silva 11503 in UC ).
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CAPE MENDOCINO						Pl													Common, in protected pools in the low intertidal and upper subtidal, on exposed rocky shores. Dawson 1965 a, p. 17. Kjeldsen 1995, p. 17.
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POINT ARENA						Pl													Common, in pools and on rocks in the low intertidal and upper subtidal in exposed habitats. Johansen 1966 a, p. 102. Kjeldsen 1995, p. 17.
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SAN FRANCISCO						Pl													Common, in pools and on rocks exposed to the surf in the low intertidal; subtidally associated with Pterygophora and in Macrocystis beds. Smith 1944, p. 138. Devinny and Kirkwood 1974, p. 102.
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