Jepson eFlora: Key page
Vascular Plants of California
Key to families | Table of families and genera

Key to Botrychium

View taxon page for Botrychium

(For a list of species in Botrychium, use the above link.)

Jepson Manual glossary definitions can be seen by moving your cursor over words underlined with dots.

1. Leaf divided into 2 nearly identical sporophores and no trophophore ..... B. paradoxum

1' Leaf divided into trophophore and sporophore (latter occasionally aborted)

2. Trophophore ternate, appearing to be divided into 3 ± equal segments due to great enlargement and dissection of   pair

3. Trophophore  or nearly so, stalk of sporophore 0.3–0.7 × trophophore ..... B. pumicola (2)

3' Trophophore clearly stalked, sporophore stalk generally >= trophophore ..... B. simplex

4. Middle pinnae narrowly attached to , slightly or not ; plants yellow-green; generally in seasonally moist meadows ..... var. compositum

4' Middle pinnae broadly attached to rachis, strongly decurrent; plants blue-green; generally in hard-water-saturated substrate ..... var. simplex (2)

2' Trophophore ; basal pinnae not disproportionately enlarged

5. Basal pinna pair pinnately dissected, midribbed; pinnae  to , broadest near base or middle ..... B. pinnatum

5' Basal pinna pair  or palmately dissected, fan-shaped, broadest at outer margin

6. Trophophore and sporophore joined well below mid-leaf, generally at ground level ..... B. simplex or B. pumicola (small or shade pls, see couplet 3 to separate the species)

6' Trophophore and sporophore joined generally near or distal to mid-leaf

7. Side margins of  pinnae ± parallel or, if cleft into segments, side margins of segments ± parallel

8. Trophophore ± sessile; sporophore stalk very short, pinnae elongate, longer than wide, not decurrent to rachis ..... B. lineare

8' Trophophore and sporophore stalk long, pinnae rhomboid, as long as wide, strongly decurrent ..... B. montanum

7' Side margins of simple pinnae converging at > 30°, or, if cleft into segments, segments wedge-shaped

9. Side margins of basal pinnae converging at (90)150–180°; pinnae touching to overlapping, especially toward tip

10. Middle pinnae , ± perpendicular to rachis; pinna texture delicate, outer margins generally finely  to ; basal sporophore branches often downturned, occurring in or near saturated substrate ..... B. crenulatum (2)

10' Middle pinnae ; pinna texture firm, outer margins entire to wavy to coarsely toothed; basal sporophore branches generally not downturned

11. Spores 33–39 µm in longest diam ..... B. lunaria

11' Spores 46–57 µm in longest diam ..... B. yaaxudakeit

9' Side margins of basal pinnae converging at 30–120(160)°; pinnae generally well-spaced to touching

12. Pinnae spreading, ± perpendicular to rachis; fertile portion of sporophore triangular to ; basal sporophore branches often downturned

13. Pinnae fan-shaped, texture delicate, deep green, outer margins generally finely crenate to dentate; occurring in or near saturated substrate ..... B. crenulatum (2) (sun plants)

13' Pinnae  or rounded without a sharp angle between side and outer margin, texture thick, yellow-green, outer margins entire or shallowly cleft; occurring in well-drained sites ..... B. tunux

12' Pinnae ascending; fertile portion of sporophore oblong to deltate; basal sporophore branches not downturned

14. Pinnae entire to symmetrically cleft into 2–4 segments, outer margins dentate; sporophore branches short, stiffly  ..... B. ascendens

14' Pinnae entire to irregularly cleft or shallowly lobed, outer margins entire to coarsely toothed or lobed; sporophore branches long, spreading ..... B. minganense


Please use this Google Form for Contact/Feedback

Citation for the whole project: Jepson Flora Project (eds.) . Jepson eFlora, [accessed on ]

Citation for an individual treatment: [Author of taxon treatment] [year]. [Taxon name] in Jepson Flora Project (eds.) Jepson eFlora, [URL for treatment]. Accessed on .

We encourage links to these pages, but the content may not be downloaded for reposting, repackaging, redistributing, or sale in any form, without written permission from The Jepson Herbarium.