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Key to HorkeliaView taxon page for Horkelia
(For a list of species in Horkelia, use the above link.) Jepson Manual glossary definitions can be seen by moving your cursor over words underlined with dots. 1. Flowers generally in head-like clusters; plant generally rosetted or tufted; NW, CaR, SN, MP, n SNE – hypanthium  Structure generally in the shape of a tube, cup, or bowl, derived from the fused lower portions of the perianth and stamens, from which these parts seem to arise, and to which the ovary wall is fused in an inferior ovary (to which the ovary wall is partially fused in a half-inferior ovary; from which the ovary is free in a superior ovary). bractlets < 0.5 mm wide; hypanthium generally 2–5 mm wide; lower pedicel  Stalk of an individual flower in an inflorescence, or the corresponding structure in fruit. often reflexed  Abruptly bent or curved downward or backward. in fruit; > 300 m 2. Leaflets 2–5 per side, entire  Having margins that are continuous and smooth (i.e., without teeth, lobes, etc.). to ± 3-toothed; petal linear  Elongate, with nearly parallel sides; narrower than elliptic or oblong. to obovate; fruit 1.5–2 mm; 300–2500 m 3. Stem densely glandular above, ascending  Curving or angling upward from base, or about 30-60 degrees less than vertical or away from axis of attachment. to erect  Upright; vertically oriented. ; petal generally 2.5–4 mm, generally widely obovate – nw KR ..... H. congesta var. nemorosa 3' Stem ± glabrous to sparsely glandular above, often decumbent  lying mostly flat on the ground but with tips curving up. ; petal 1.5–4 mm, linear to widely oblanceolate ..... H. tridentata 4. Petal ± widely oblanceolate, often < sepal; hypanthium 2.5–5 mm wide, inner wall hairy (except in NCoRH); KR, NCoR, n SNH ..... var. flavescens 4' Petal generally narrow-oblanceolate, generally >= sepal; hypanthium generally 2–3.5 mm wide, inner wall generally glabrous; e KR, CaR, SN, MP ..... var. tridentata 2' Leaflets 3–15 per side, > 3-toothed; petal ± wedge-shaped; fruit 1–1.8 mm; > 950 m ..... H. fusca 5. Leaflets generally divided ± 3/4 to base, generally 8–15 per side, ± crowded; filament generally < 0.5 mm, generally wider than long – s CaRH ..... var. tenella (2) 5' Leaflets generally toothed <= 1/2 to base, generally 4–8 per side, ± separated; filament often > 0.5 mm, generally longer than wide 6. Petal 4–6 mm; leaflets generally 10–20 mm, toothed < 1/4 to base, generally ± obovate; anther ± 0.6 mm; style 1–1.5 mm; fruit ± 1.8 mm; Wrn ..... var. pseudocapitata 6' Petal 2–4 mm; leaflets generally 5–15 mm, toothed 1/4–1/2 to base, ± round to wedge-shaped; anther ± 0.4 mm; style ± 1 mm; fruit 1–1.2 mm; n California (except Wrn), SNH 1' Flowers generally either in ± flat-topped clusters or inflorescence open; plant often matted; CA-FP, W&I 8. Pedicel ± recurved  Gradually curved downward or backward. in fruit; inflorescence open, often ± 1-sided 9. Petal 4–7 mm; leaflets toothed 1/4 to midrib; pistils generally 20–50; n&c SNF (especially Ione Formation) ..... H. parryi 8' Pedicel erect in fruit; inflorescence open or dense, not 1-sided 10' Filament >= 0.5 mm, generally longer than wide; style 1–4 mm 11. Plant generally rosetted from simple  Composed of a single part; undivided; unbranched. or few-branched caudex  Generally short, sometimes woody, more or less vertical stem of a perennial, at or beneath ground level. ; resinous odor weak; leaflets generally < 10-lobed; fruit >= 2 mm; often ± serpentine clay 12. Leaflets 5–10 per side, 5–25 mm, lobed > 3/4 to base; stem base hairs 2–3 mm; inflorescence open to dense; hypanthium 3.5–5 mm wide; petal ± obovate, cream; pistils 5–15; style 2.5–4 mm ..... H. daucifolia 13. Leaflets ± gray, lobes  1. A major expansion or bulge, such as on the margin of a leaf, sepal, or petal, or on the surface of an ovary. 2. The free tips of otherwise fused structures, such as sepals or petals; larger than teeth. 2–5, 1–3 mm wide; petal 2–4 mm wide; pedicels Stalk of an individual flower in an inflorescence, or the corresponding structure in fruit. nonglandular ..... var. daucifolia 12' Leaflets ± 15 per side, 2–10 mm, generally lobed ± 1/2 to base; stem base hairs ± 1 mm; inflorescence open; hypanthium 2–3 mm wide; petal obcordate, often pink-tinged; pistils 2–6; style ± 1.5–2 mm 14. Stipules of basal  At or near the base of a plant or plant part. Especially said of leaves clustered near the ground or of a placenta confined to the base of an ovary. leaves pinnately divided; leaves ± green, 5–15 cm ..... H. howellii 14' Stipules of basal leaves entire or forked  Pertaining to a hair or other structure that branches into two parts. ; leaves silvery, 3–10 cm ..... H. sericata 11' Plant generally tufted to matted from a few- to many-branched caudex; resinous odor strong; leaflets often > 10-toothed or -lobed; fruit often < 2 mm; generally non-serpentine 15. Leaflets pinnately veined, oblong  Longer than wide, with nearly parallel sides; wider than linear. to obovate, ± evenly > 10-toothed – hypanthium width 4–8 mm, > 2 × length, bractlets ± 2–3 mm wide; CW, SW 16. Leaflets 1–3 per side, < distinct terminal leaflet  A leaf-like unit of a compound leaf; distinguished from a leaf by the absence in its axil of a bud, branch, thorn, or flower; lacking lateral, basal appendages (stipules); either simple (leaf 1-compound, with primary leaflets) or compound (leaf 2-compound, with primary and secondary leaflets; 3-compound, with primary, secondary, and tertiary leaflets, etc). , teeth generally double, generally > 20; petal ± round, ± 5 mm wide; 400–1300 m; PR ..... H. truncata 16' Leaflets 5–12 per side, ± = generally indistinct terminal leaflet, teeth generally single, ± 10–15; petal obovate or narrower, 1.5–4 mm wide; < 870 m; CCo, SCoRO, SCo ..... H. cuneata 17. Hairs ± all glandular; inflorescence open; hypanthium inner rim ± glabrous; filament base generally 0.5–2 mm wide ..... var. puberula 17' Hairs mostly glandless; inflorescence open to dense; hypanthium inner rim generally hairy; filament base 0.5–1 mm wide 18. Inflorescence generally ± open; hairs ± spreading  Oriented more or less perpendicularly to the axis of attachment; often, more or less horizontal. , ± dense; glands ± obvious ..... var. cuneata 18' Inflorescence dense to ± open; hairs ascending to appressed  Parallel or nearly parallel to and often in contact with surface of origin; used to describe the disposition of hairs, leaves, pedicels, etc. , dense; glands hidden, if any ..... var. sericea 15' Leaflets ± palmately veined at base, wedge-shaped or ± round-ovate, ± unevenly few- to many-toothed or -lobed 19. Hypanthium width 4–10 mm, 1–2 × length, bractlets ovate  Egg-shaped (i.e., widest below the middle) in two dimensions (i.e., in one plane), as a leaf. , generally ± 2 mm wide, ± = sepal; leaflets round to ovate, 5–60 mm, generally ± separated, generally > 20-toothed or -lobed; stem ± leafy; fruit ± 1 mm ..... H. californica 20. Leaflets 3–5 per side, generally unlobed, 15–60 mm, elliptic  In the shape of a flattened circle or ellipse; wider than linear to ovate – hypanthium inner wall glabrous; sepal not red-mottled; style 2–3 mm; CW ..... var. frondosa 20' Leaflets 4–9 per side, ± lobed, 5–40 mm, ovate to round 21. Leaflets generally few-lobed ± 1/2 to base, 10–40 mm; sepal red-mottled inside; hypanthium bractlets generally toothed; hypanthium inner wall ± hairy; filament generally 1.5–3 mm; style generally 3–4 mm; NCo, CCo ..... var. californica 21' Leaflets lobed 1/2–3/4 to base, 5–25 mm; sepal not red-mottled; hypanthium bractlets generally entire; hypanthium inner wall glabrous; filament generally 0.5–1.5 mm; style generally 2–3 mm; NCoR, c&s SN, SnFrB, SnBr ..... var. elata 19' Hypanthium width generally < 6 mm, generally 2+ × length, bractlets lanceolate  Narrowly elongate, widest in the basal half, often tapered to an acute tip. to ovate, <= 2 mm wide, < sepal; leaflets round to wedge-shaped, generally < 15 mm, often crowded, < 20-toothed or -lobed; stem leaves few, ± reduced; fruit > 1 mm 22. Stem < 25 cm; hypanthium bractlets generally < 0.5 mm wide; leaves 2–10 cm; fruit 1.5–2.5 mm; > 2000 m; KR, s SNH, W&I 23. Hypanthium densely hairy on inner rim; sepal 3.5–6 mm, 2+ × hypanthium length; longer hairs ± 1 mm, soft – ne KR ..... H. hendersonii 23' Hypanthium ± hairy on inner wall; sepal generally 2–4 mm, < 2 × hypanthium length; longer hairs ± 0.5 mm, stiff 22' Stem 10–70 cm; hypanthium bractlets generally >= 0.5 mm wide; leaves 3–20 cm; fruit 1.5–2 mm; < 2800 m; NCo, NCoR, CW, SW 25. Stem decumbent to ascending, generally 10–30 cm; plant matted, odor strong; inflorescence ± dense; leaves generally 4–10 cm; leaflets generally 5–10 per side, ± crowded – hairs dense; NCo, CCo ..... H. marinensis 25' Stem ascending to erect, 10–70 cm; plant tufted or matted, odor strong or not; inflorescence generally ± open; leaves generally 3–20 cm; leaflets 6–16 per side, separated or crowded 26. Hypanthium bractlets 1–2 mm wide, lanceolate to ovate; anther ± 1 mm – hypanthium generally 3–6 mm wide; stem hairs spreading; SCoRO ..... H. yadonii 26' Hypanthium bractlets generally 0.5–1 mm wide, linear or elliptic to lanceolate; anther 0.5–1 mm 27. Stem and petiole  Leaf stalk, connecting leaf blade to stem; sometimes more or less indistinct. hairs generally spreading; plant often green 28' Leaflets divided > 1/2 to base, lobes linear to oblanceolate; hypanthium inner wall ± hairy; NCo, NCoRO, SnFrB ..... H. tenuiloba 27' Stem and petiole hairs generally ascending to appressed; plant generally ± gray – hypanthium inner wall ± hairy
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