Jepson eFlora: Taxon page
Vascular Plants of California
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Thelypteris puberula var. sonorensis

Higher Taxonomy
Family: ThelypteridaceaeView Description 
Habit: Plants generally in soil; rhizome short- to long-creeping, prostrate to erect, [trunk-like], scales generally hairy, +- brown. Leaf: stipe ×-section at base with 2 crescent-shaped vascular strands fusing into 1 U-shaped strand distally; blade 1[2]-pinnate, generally with needle-like, clear hairs on axes and/or between veins; rachis, costae generally grooved adaxially except between axes; veins free or in regular, net-like pattern. Sporangia: sori on veins, generally round; indusia reniform or round-reniform [0]; stalk cells in 3 rows; spores generally elliptic, scar generally linear.
Genera In Family: 1 genus (5--32 genera), +- 1000 species: worldwide, especially tropics; several cultivated.
eFlora Treatment Author: John C. Game, Alan R. Smith & Thomas Lemieux
Scientific Editor: Alan R. Smith, Thomas J. Rosatti.
Genus: ThelypterisView DescriptionDichotomous Key

Leaf: pinnae entire to generally deeply lobed, veins generally not forked.
Etymology: (Greek: female fern; ancient name for delicate fern)
Reference: Smith & Cranfill 2002 Amer Fern J 92:131--149
Unabridged Reference: Smith 1971 Univ Calif Publ Bot 59:1--143; Smith, A. R. 1990. Thelypteridaceae. Pp. 263--272 in The families and genera of vascular plants, ed. K. Kubitzki. Vol. 1. Pteridophytes and gymnosperms, ed. K. Kramer & P. Green. Berlin: Springer-Verlag
Thelypteris puberula (Baker) C.V. Morton var. sonorensis A.R. Sm.
Habit: Rhizome creeping, 3--8 mm wide; evergreen. Leaf: +- regularly spaced, generally 50--120 cm, 15--30 cm wide; stipe scales lanceolate, brown, not persistent; blade thick, abaxially +- nonglandular, +- densely hairy on axes, veins, and between veins; pinnae deeply lobed, lobes narrowly oblong, entire. Sporangia: sori small, round; indusia densely hairy.
Ecology: Along streams, seepage areas; Elevation: 50--800 m. Bioregional Distribution: SCo, WTR, SnGb, SnJt; Distribution Outside California: to Arizona, southern Mexico. Note: Rarely cultivated in southern California. 1 other variety, Thelypteris puberula var. puberula, Mexico to Costa Rica.
Unabridged Note: Christella puberula A. Löve & D. Löve, published [1977 Taxon 26:326] for the nominate element of Thelypteris puberula, may reflect a superior classification, but a combination that would include var. sonorensis within Christella puberula has not been made. Recent molecular evidence [A.R. Smith & R.B. Cranfill 2002 Taxon 56:1037--1050] suggests that Christella itself may be polyphyletic and possibly not separable from Cyclosorus s.l. Some combinations of supposed Christella taxa within Cyclosorus, including "Cyclosorus puberulus," have not yet been made.
Jepson eFlora Author: John C. Game, Alan R. Smith & Thomas Lemieux
Reference: Smith & Cranfill 2002 Amer Fern J 92:131--149
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)
Listed on CNPS Rare Plant Inventory

Previous taxon: Thelypteris nevadensis
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Botanical illustration including Thelypteris puberula var. sonorensis

botanical illustration including Thelypteris puberula var. sonorensis


Citation for this treatment: John C. Game, Alan R. Smith & Thomas Lemieux 2012, Thelypteris puberula var. sonorensis, in Jepson Flora Project (eds.) Jepson eFlora, /eflora/eflora_display.php?tid=67105, accessed on May 12, 2021.

Citation for the whole project: Jepson Flora Project (eds.) 2021, Jepson eFlora,, accessed on May 12, 2021.

No expert verified images found for Thelypteris puberula var. sonorensis.

Geographic subdivisions for Thelypteris puberula var. sonorensis:
SCo, WTR, SnGb, SnJt
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).


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Data provided by the participants of the  Consortium of California Herbaria.
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All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.

CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).