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Vascular Plants of California
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Nemacladus rigidus


Higher Taxonomy
Family: CampanulaceaeView DescriptionDichotomous Key
Common Name: BELLFLOWER FAMILY
Habit: Annual to perennial herb [tree]. Leaf: generally cauline, generally simple, generally alternate, petioled or not; stipules 0. Inflorescence: cyme, panicle, raceme, spike, or flowers 1; terminal or in axils of leaf-like or reduced bracts. Flower: bisexual, cleistogamous or open, radial or bilateral, inverted (pedicel twisted 180°) or not; hypanthium generally present, +- fused to ovary; sepals generally 5; corolla radial to 2-lipped, petals generally fused, tube deeply divided on 1 side or not, lobes generally 5; stamens 5, free or +- fused (anthers, filaments fused into tube or filaments fused above middle); ovary inferior or 1/2 inferior (superior in fruit), chambers 1--3, placentas axile or parietal, ovules many, style generally 1, 2--5-branched. Fruit: generally capsule, open on sides or top by pores or short valves. Seed: many.
Genera In Family: +- 90 genera, +- 2500 species: worldwide. Note: Some cultivated for ornament (Campanula, Jasione, Lobelia). Subfamilies sometimes treated as families. Positions of flower parts given after flowering inversion, if any. Parishella moved to Nemacladus.
eFlora Treatment Author: Nancy R. Morin, except as noted
Scientific Editor: Thomas J. Rosatti.
Genus: NemacladusView DescriptionDichotomous Key


Habit: Annual, from taproot. Stem: prostrate, decumbent, or erect; base generally +- brown or +- purple; branches 0 or below middle. Leaf: basal; petiole short or 0. Inflorescence: +- raceme-like; bract 1 per flower, small; pedicel generally thread-like. Flower: inverted or not; sepals linear to triangular; corolla +- radial or 2-lipped, lobes 5; filaments free at base, fused into tube around style distally, appendages attached to a stalk or directly on 2 adjacent filaments, each with 2--12 cells, anthers free, all alike; ovary superior to 1/2 inferior, hemispheric to obconic, nectary glands 3, mounded or donut-like, on free part of ovary, stigma 2-lobed, papillate. Fruit: generally > hypanthium, hemispheric to fusiform, top pointed or rounded, chambers 2; open at top generally by 2 valves (or circumscissile). Seed: elliptic to oblong.
Etymology: (Greek: thread-like branch) Note: In descriptions, "filaments" including both free and fused parts thereof.
Unabridged Note: Taxonomic changes from TJM (1993) based on ITS, atpB, morphology. Parishella californica nested within Nemacladus, in which it is here treated.
Reference: Morin 2008 J Bot Res Inst Texas 2:397--400
Unabridged Reference: McVaugh 1942 N Amer Flora 32A:1--134
Nemacladus rigidus Curran
NATIVE
Habit: Spreading to decumbent, 2--4(9) cm, branched from base. Stem: spreading, stout; base shiny, purple. Leaf: 5--10 mm, elliptic to oblanceolate, fleshy, entire to scalloped, hairy, narrowed to wide petiole. Inflorescence: axis strongly zigzag; bracts 2--3 mm, widely elliptic, spreading or reflexed; pedicels 5--12 mm, 0.2 mm diam, spreading, straight, in age curved. Flower: not inverted; hypanthium 1 mm; 2 adaxial, middle abaxial sepals linear-elliptic, 0.7 mm, 2 flanking abaxial sepals widely triangular, 1.4 mm; corolla divided to base, 2-lipped, white, 3 adaxial lobes erect, 0.2--0.4 mm, ovate, glabrous, veins and tips maroon, 2 abaxial 0.4--0.5 mm, oblong, acute; filaments declined, 1.2--1.6 mm, tip +- curved, glabrous, appendage pad near base of filaments, cells wide, blunt, anthers 0.2--0.3 mm; ovary +- inferior in fruit. Fruit: 3--4 mm, base oblique, tip pointed. Seed: 0.6--0.7 mm, elliptic, with wide zigzag ridges alternate pitted rows.
Ecology: Bare soil, sand; Elevation: 200--2500 m. Bioregional Distribution: GB; Distribution Outside California: eastern Oregon, Idaho, Nevada. Flowering Time: May--Jun
Jepson eFlora Author: Nancy R. Morin
Reference: Morin 2008 J Bot Res Inst Texas 2:397--400
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)

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botanical illustration including Nemacladus rigidus

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Citation for this treatment: Nancy R. Morin 2012, Nemacladus rigidus, in Jepson Flora Project (eds.) Jepson eFlora, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=34511, accessed on April 19, 2024.

Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on April 19, 2024.

Nemacladus rigidus
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©2010 Steve Matson
Nemacladus rigidus
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©2010 James Morefield
Nemacladus rigidus
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©2010 Steve Matson
Nemacladus rigidus
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©2010 Steve Matson
Nemacladus rigidus
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©2010 Steve Matson

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Geographic subdivisions for Nemacladus rigidus:
GB
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).





 

Data provided by the participants of the  Consortium of California Herbaria.
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Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).