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Vascular Plants of California
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Calycadenia multiglandulosa

Higher Taxonomy
Family: Asteraceae (Compositae)View DescriptionDichotomous Key
Habit: Annual to tree. Leaf: basal and/or cauline, alternate, opposite, rarely whorled, simple to 2+ × compound. Inflorescence: 1° inflorescence a head, resembling a flower, of several types (see below), 1--many in generally +- cyme-like cluster; each head generally with +- calyx-like involucre of 1--many series of phyllaries (involucre bracts); receptacle of head flat to conic or columnar, paleate (bearing paleae = receptacle bracts) or epaleate; flowers 1--many per head. Flower: bisexual, unisexual, or sterile, +- small, of several types (see below); calyx 0 or modified into +- persistent pappus of bristles, scales, and/or awns; corolla radial or bilateral (0), lobes generally (0)3--5; stamens 4--5, filaments generally free, generally fused to corolla at tube/throat junction, anthers generally fused into cylinder around style, anther base generally rounded or cordate (deeply sagittate or with tail-like appendages), tip (= flattened appendage) generally projecting beyond pollen sac; pistil 1, 2-carpeled, ovary inferior, 1-chambered, 1-seeded, placenta basal, style 1, tip generally +- 2-branched (except in some staminate disk flowers), branch tips truncate or generally bearing +- brush-like appendages; stigmas 2, generally on adaxial faces of style branches. Fruit: achene (also called a cypsela) (drupe in Chrysanthemoides), cylindric to ovoid, sometimes compressed, generally deciduous with pappus attached.
Genera In Family: +- 1500 genera, 23000 species: worldwide, many habitats. Note: Flower and head types differ in form and sexual condition. A disk flower has a generally radial corolla, with a cylindric tube, expanded throat, and generally 5 lobes. Disk flowers are generally bisexual and fertile but occasionally staminate with reduced ovaries. Discoid heads comprise only disk flowers. A radiant head is a variant of a discoid head, with peripheral disk flower corollas expanded, often bilateral. A ray flower corolla is bilateral, generally with a slender tube and flattened petal-like ray (single lip composed of generally 3 lobes). Ray flowers are generally pistillate or sterile (occasionally lacking styles). Radiate heads have peripheral ray flowers and central disk flowers. Disciform heads superficially resemble discoid heads, with pistillate or sterile flowers that lack rays, together with or separate from disk flowers. A ligulate flower is bisexual, with a bilateral, generally ephemeral corolla and 5-lobed ligule. Liguliflorous heads comprise only ligulate flowers. See glossary p. 31 for illustrations of family characteristics. Echinops sphaerocephalus L., Gaillardia aristata Pursh, Gaillardia pulchella Foug., Hymenothrix loomisii S.F. Blake, Tagetes erecta L., Thelesperma megapotamicum (Spreng.) Kuntze are waifs. Melampodium perfoliatum Kunth, historic urban waif. Ageratum conyzoides L., Guizotia abyssinica (L. f.) Cass., Santolina chamaecyparisus L., orth. var. are rare or uncommon escapes from cultivation. Dyssodia papposa, Ismelia carinata (Schousb.) Sch. Bip. [Chrysanthemum carinatum Schousb.], Mantisalca salmantica (L.) Briq. & Cavill. are historical or extirpated waifs in California. Inula helenium L. not documented in California. Taxa of Aster in TJM (1993) treated here in Almutaster, Doellingeria, Eurybia, Ionactis, Oreostemma, Sericocarpus, Symphyotrichum; Chamomilla in Matricaria; Bahia in Hymenothrix; Cnicus in Centaurea; Conyza in Erigeron and Laennecia; Dugaldia in Hymenoxys; Erechtites in Senecio; Hymenoclea in Ambrosia; Lembertia in Monolopia; Osteospermum ecklonis in Dimorphotheca; Picris echioides in Helminthotheca; Prionopsis in Grindelia; Raillardiopsis in Anisocarpus and Carlquistia; Schkuhria multiflora in Picradeniopsis; Trimorpha in Erigeron; Venidium in Arctotis; Whitneya in Arnica. Amauriopsis in TJM2 (2012) treated here in Hymenothrix; Arida in Leucosyris; Bahia in Picradeniopsis; Eucephalus in Doellingeria.
Unabridged Note: Largest family of vascular plants in California and of eudicots globally.
eFlora Treatment Author: David J. Keil, except as noted
Scientific Editor: David J. Keil, Bruce G. Baldwin.
Genus: CalycadeniaView DescriptionDichotomous Key

Habit: Annual, erect to spreading, with coarse tack-shaped glands, and often simple glandular hairs, aromatic; generally self-sterile. Leaf: basal +- many, mostly deciduous by flowering, occasionally persistent, margins often curled under, and cauline, generally alternate and reduced distally on stem, +- sessile, +- linear, entire or sparsely toothed, +- scabrous, generally with longer stiff hairs, especially on proximal margins. Inflorescence: heads radiate, axillary, often +- sessile, and terminal, 1 or clustered; peduncle bracts linear or narrowly lanceolate to widely oblanceolate, margins thickened, rounded, defining a +- prominent central abaxial channel, especially proximally, +- bristly-ciliate, especially proximally, some or all with 1+ tack-like glands; phyllaries +- equal to shorter than paleae, each partly enfolding a ray fruit; receptacle +- flat, hairy; paleae in one series between ray and disk flowers, +- fused, forming a receptacle cup, phyllaries, paleae generally with abaxial simple and/or tack-like glands and appressed and/or spreading hairs. Ray Flower: 1--6; ray 2--12 mm, 3-lobed, white to rose or yellow, occasionally with red spot near base. Disk Flower: 1--25; corolla 2.5--10 mm, colored +- like rays; anthers generally purple, tips ovate to triangular; style branches long, bristly. Fruit: ray fruit 1.5--4 mm, +- 3-angled, adaxial face +- flat, abaxial face rounded to +- angled, pappus 0; disk fruit 2--4 mm, angled, base tapered, +- appressed-hairy to glabrous, pappus 0 or generally of 6--16 scales, all +- lanceolate or some blunt and shorter.
Species In Genus: +- 10 species: especially California (Oregon). Etymology: (Greek: cup gland)
eFlora Treatment Author: Robert L. Carr & Gerald D. Carr
Reference: Carr & Carr 2006 FNANM 21:270--276
Calycadenia multiglandulosa DC.
Stem: 1--7 dm, +- simple or branched, often with diffuse, relatively short, many-bracted branchlets, generally hairy, glandular, especially distally. Leaf: generally alternate, occasionally opposite to mid-stem or beyond, proximal often persistent, 3--8 cm. Inflorescence: spike-like to highly congested, especially distally; peduncle bracts 4--20 mm, often +- red, linear to narrowly lanceolate, occasionally narrowed and +- cylindric toward tip, tack-like glands generally 2--6+; phyllaries 4--8.5 mm, paleae 4.5--10 mm, both abaxially often long-hairy, glandular, +- red, tack-like glands (0)1--15+. Ray Flower: 2--6; ray 5--10 mm, white or cream to rose or yellow; central lobe << and narrower than lateral lobes, widest at base, symmetric, laterals strongly asymmetric, sinuses generally 1/3--1/2 ray. Disk Flower: 4--20, corolla 6--10 mm. Fruit: ray fruit smooth, glabrous to +- appressed-hairy; disk pappus scales +- 11, lanceolate, acuminate, 2--3 shorter, blunt. Chromosomes: 2n=12.
Ecology: Common. Generally dry, open valleys, hillsides, rocky ridges; Elevation: 50--1100 m. Bioregional Distribution: NCoR, SN, GV, SnFrB, SCoRI. Flowering Time: May--Oct Note: Variable.
Synonyms: Calycadenia hispida (Greene) Greene; Calycadenia multiglandulosa subsp. bicolor (Greene) D.D. Keck; Calycadenia multiglandulosa subsp. cephalotes (DC.) D.D. Keck; Calycadenia hispida subsp. reducta D.D. Keck; Calycadenia multiglandulosa subsp. robusta D.D. Keck; Calycadenia multiglandulosa subsp. cephalotes (DC.) D.D. Keck
Jepson eFlora Author: Robert L. Carr & Gerald D. Carr
Reference: Carr & Carr 2006 FNANM 21:270--276
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)

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Botanical illustration including Calycadenia multiglandulosa

botanical illustration including Calycadenia multiglandulosa


Citation for this treatment: Robert L. Carr & Gerald D. Carr 2012, Calycadenia multiglandulosa, in Jepson Flora Project (eds.) Jepson eFlora, /eflora/eflora_display.php?tid=1875, accessed on June 13, 2021.

Citation for the whole project: Jepson Flora Project (eds.) 2021, Jepson eFlora,, accessed on June 13, 2021.

Calycadenia multiglandulosa
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© 2009 Neal Kramer
Calycadenia multiglandulosa
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© 2008 Neal Kramer
Calycadenia multiglandulosa
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© 2016 Neal Kramer
Calycadenia multiglandulosa
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© 2009 Neal Kramer
Calycadenia multiglandulosa
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© 2007 George W. Hartwell
Calycadenia multiglandulosa
click for enlargement
© 2009 Neal Kramer

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map of distribution 1
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Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).