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Vascular Plants of California
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Johnstonella angelica

Higher Taxonomy
Family: BoraginaceaeView DescriptionDichotomous Key

Habit: Annual, perennial herb, or shrub, often bristly or sharp-hairy. Stem: prostrate-decumbent to erect. Leaf: basal and/or cauline, simple, generally alternate, sometimes opposite, especially at base. Inflorescence: cymes, arranged singly or in groups of 2--5, generally coiled in flower, generally elongating in fruit. Flower: bisexual, generally radial; sepals 5, free or fused at least at base; corolla 5-lobed, salverform, funnel-shaped, rotate, or bell-shaped, appendages (often called "fornices") 0 or 5 at top of tube, when present often differentially pigmented, alternate stamens; stamens epipetalous; ovary superior, 4-lobed, style 1, entire or minutely 2-lobed (2-branched). Fruit: nutlets 1--4, when > 1, all similar (often called "homomorphic") or 1 or 2 dissimilar in size and/or shape from the others (often called "heteromorphic"), free (fused), smooth to roughened, prickly or bristly or not.
Genera In Family: +- 90 genera, +- 1600--1700 species: mostly temperate, especially western North America, Mediterranean; some cultivated (Borago, Echium, Myosotis, Symphytum). Toxicity: Many genera may be TOXIC from pyrrolizidine alkaloids or accumulated nitrates. Note: Sometimes still treated in broader sense of TJM2 (e.g., APG IV 2016 Bot J Linn Soc 181:1--20), but recent evidence (Luebert et al. 2016) supports segregation, for our flora, of the families Ehretiaceae, Heliotropiaceae, Hydrophyllaceae, Lennoaceae, and Namaceae.
eFlora Treatment Author: Michael G. Simpson, C. Matt Guilliams, Kristen Hasenstab-Lehman & Ronald B. Kelley
Scientific Editor: Bruce G. Baldwin, C. Matt Guilliams, Kristen Hasenstab-Lehman, David J. Keil, Ronald B. Kelley, Robert W. Patterson, Thomas J. Rosatti & Michael G. Simpson
Johnstonella angelica (I.M. Johnst.) Hasenstab & M.G. Simpson
Habit: Annual 15--25 cm. Stem: branches from base, lateral branches numerous; smooth proximally, canescent-strigose distally. Leaf: 1--2.5 cm, linear, strigose, hairs bulbous-based, especially abaxially. Inflorescence: in 1--3s, dense; bracts 0; pedicel not elongated in fruit, <= 0.5 mm. Flower: calyx 2--2.5 mm in fruit, ascending, lobes linear-lanceolate, midvein thickened, soft-bristly; corolla limb < 1 mm diam. Fruit: nutlets 4, larger, persistent nutlet +- 0.9--1.2 mm, three smaller nutlets +- 0.6--0.8 mm, all lance-ovate, +- brown, pale-tubercled, margin sharp-angled, base rounded; abaxially shallowly convex, ridge 0; adaxially bi-convex, attachment scar edges not raised, abutted near tip, narrowly triangular-gapped in lower 1/3. Chromosomes: 2n=18.
Ecology: Desert wash, succulent scrub; Elevation: 210 m. Bioregional Distribution: DSon (Borrego Springs, San Diego Co.); Distribution Outside California: Baja California, Baja California Sur, Sonora. Flowering Time: Likely Feb--Apr (almost year-round in parts of Mexico, depending on monsoonal rains) Note: Previously known only from Baja California and Baja California Sur. A recent collection in San Diego Co. (Simpson et al. 2020) thought to possibly be a waif, but now known to be well-established; full North American distribution awaits further surveys. Elevational range 5--700 m in Mexico.
Synonyms: Cryptantha angelica I.M. Johnst.
Jepson eFlora Author: Michael G. Simpson, Kristen E. Hasenstab-Lehman & Ronald B. Kelley
Reference: Johnston 1925 Contr Gray Herbarium 74:1--125; Simpson & Hasenstab 2009 Crossosoma 35:1--59; Hasenstab-Lehman & Simpson 2012 Syst Bot 37:738--757; Simpson et al. 2014 Taxon 63:930--931; Simpson et al. 2017 Taxon 66:1406--1420; Simpson et al. 2019 Phytotaxa 425:279--289; Simpson et al. 2020 Madroño 67:5--8.
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Citation for this treatment: Michael G. Simpson, Kristen E. Hasenstab-Lehman & Ronald B. Kelley 2021, Johnstonella angelica, in Jepson Flora Project (eds.) Jepson eFlora, Revision 9,, accessed on October 02, 2023.

Citation for the whole project: Jepson Flora Project (eds.) 2023, Jepson eFlora,, accessed on October 02, 2023.

Johnstonella angelica
click for enlargement
© 2022 Michael G. Simpson
Johnstonella angelica
click for enlargement
© 2022 Michael G. Simpson
Johnstonella angelica
click for enlargement
© 2022 Michael G. Simpson

More photos of Johnstonella angelica in CalPhotos

Geographic subdivisions for Johnstonella angelica:
DSon (Borrego Springs, San Diego Co.)
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).


Data provided by the participants of the  Consortium of California Herbaria.
View all CCH records
All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.

CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).