Habit: Shrub, generally rhizomed.
Stem: spreading to erect, branched, spiny or not, vine-like or not; inner bark, wood generally bright yellow; over-wintering bud scales deciduous or not.
Leaf: simple or pinnately compound, cauline, alternate, generally leathery, generally persistent; leaflets generally 3--11, +- round to lanceolate, generally spine-toothed.
Inflorescence: raceme, axillary or terminal.
Flower: sepals 9 in 3 whorls of 3; petals 6 in 2 whorls of 3, bases generally glandular; stamens 6; ovules 2--9, stigma +- spheric.
Fruit: berry, spheric to elliptic, generally purple-black.
Species In Genus: +- 600 species: temperate worldwide.
Etymology: (Latin: ancient Arabic name for barberry)
Toxicity: Roots often TOXIC: spines may inject fungal spores into skin.
Note: Contact with filament causes stamen to snap inward, possibly to deposit pollen on pollinator.
Jepson eFlora Author: Michael P. Williams
Reference: Kim 2004 J Pl Res 117:175--182
Unabridged Reference: Moran 1982 Phytologia 52:221--226, for relationship between Berberis, Mahonia.Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)Key to Berberis
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