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Vascular Plants of California
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Eremogone macradenia var. macradenia

Higher Taxonomy
Family: CaryophyllaceaeView DescriptionDichotomous Key
Common Name: PINK FAMILY
Habit: Annual to perennial herb; rarely dioecious (Silene), taprooted or rhizome generally slender. Leaf: simple, generally opposite (subwhorled), entire, pairs at nodes often +- connected at bases; stipules generally 0; petiole generally 0. Inflorescence: generally cyme, generally open; flowers 1--many; involucre generally 0 (present in most Dianthus, Petrorhagia). Flower: generally bisexual, radial; hypanthium often present but obscure; sepals (4)5, +- free or fused into a tube, margins generally scarious, more so on inner 2 or not, tube generally not scarious, awns generally 0; petals (4)5 or 0, generally tapered to base (or with claw long, limb expanded), entire to 2--several-lobed, limb generally without scale-like appendages adaxially, generally without ear-like lobes at base; stamens generally 10, generally fertile, generally free, generally from ovary base; nectaries 0 or 5; ovary superior, generally 1-chambered, placentas basal or free-central, styles 2--5 with 0 branches or 1 with 2--3 branches. Fruit: capsule or utricle (rarely +- dehiscent), generally sessile. Seed: appendage generally 0 (present in Moehringia).
Genera In Family: +- 100 genera, 3000 species: widespread, especially arctic, alpine, temperate northern hemisphere; some cultivated (Agrostemma, Arenaria, Atocion, Cerastium, Dianthus, Gypsophila, Lychnis, Sagina, Saponaria, Silene). Note: Apetalous Caryophyllaceae can also be keyed in Rabeler & Hartman 2005 FNANM 5:5--8. Taxa of Minuartia in TJM2 treated here in Cherleria and Sabulina; Pseudostellaria in Hartmaniella and Torreyostellaria; Vaccaria in Gypsophila; Velezia in Dianthus.
eFlora Treatment Author: Ronald L. Hartman (deceased) & Richard K. Rabeler, except as noted
Scientific Editor: Bruce G. Baldwin & Thomas J. Rosatti.
Genus: EremogoneView DescriptionDichotomous Key

Common Name: SANDWORT
Habit: Perennial herb, prostrate (non-flowering stems) or ascending to erect to mat-forming, taprooted. Leaf: needle-like to narrowly linear; vein 1. Inflorescence: terminal, open to head- or umbel-like; flowers 1--many; peduncles, pedicels 0--55 mm. Flower: hypanthium present; sepals 5, +- free, 3--7.2 mm, lance-linear to ovate, glabrous to glandular-hairy; petals 5, 2--18 mm, entire or +- notched; stamens on hypanthium; ovary +- superior, styles 3, 2.5--3 mm. Fruit: capsule, ovoid to urn-shaped; teeth 6, ascending to recurved. Seed: 1--9, +- gray, dark brown, red-brown, yellow-tan, black-purple, or +- black.
Etymology: (Greek: solitary or deserted + seed, allusion uncertain) Note: Based in part on molecular evidence (Harbaugh et al. 2010 Intl J Plant Sci 171:185--198), 2 subgenera of Arenaria treated here as Eremogone.
eFlora Treatment Author: Ronald L. Hartman (deceased) & Richard K. Rabeler
Reference: Hartman & Rabeler 2004 Sida 21:237--241
Unabridged Reference: Hartman, Rabeler, & Utech 2005 FNANM 5:56--70
Species: Eremogone macradeniaView Description 

Habit: Perennial herb, tufted, green. Stem: 20--40 cm, rounded, +- dull, glandular-hairy or not. Leaf: 20--60 mm, 0.5--2 mm wide, blunt to sharp-pointed, vein 1. Inflorescence: terminal, generally open; flowers several to many; pedicels 3--55 mm. Flower: sepals 4.5--7.2 mm, in fruit < 8 mm, acute to acuminate; petals 6--11 mm; nectaries rectangular, 2-lobed or truncate, 0.7--1.5 mm. Fruit: 6--7 mm. Seed: 4--9, 1.8--2.7 mm, +- spheric to ovate, compressed, red-brown to +- black; tubercles low, rounded to conic.

Eremogone macradenia (S. Watson) Ikonn. var. macradenia
Leaf: +- ascending, 0.8--1.2 mm wide. Inflorescence: branches (and sepals) +- glabrous. Flower: sepals 4.5--7.2 mm, in fruit < 8 mm.
Ecology: Open woodland, sagebrush flats, dry rocky slopes, alluvial deposits, often on carbonates; Elevation: 1100--2200 m. Bioregional Distribution: s SN, SnGb, SnBr, SNE, DMoj; Distribution Outside California: to Utah, Arizona. Flowering Time: Apr--Jun
Synonyms: Arenaria macradenia S. Watson var. macradenia; Arenaria macradenia var. parishiorum B.L. Rob.
Jepson eFlora Author: Ronald L. Hartman (deceased) & Richard K. Rabeler
Reference: Hartman & Rabeler 2004 Sida 21:237--241
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)

Previous taxon: Eremogone macradenia var. arcuifolia
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Botanical illustration including Eremogone macradenia var. macradeniabotanical illustration including Eremogone macradenia var. macradenia

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Citation for this treatment: Ronald L. Hartman (deceased) & Richard K. Rabeler 2012, Eremogone macradenia var. macradenia, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on July 23, 2024.

Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora,, accessed on July 23, 2024.

Eremogone macradenia  
var. macradenia
click for image enlargement
©2009 Thomas Stoughton
Eremogone macradenia  
var. macradenia
click for image enlargement
©2011 Steve Matson
Eremogone macradenia  
var. macradenia
click for image enlargement
©2011 Neal Kramer
Eremogone macradenia  
var. macradenia
click for image enlargement
©2016 James Morefield
Eremogone macradenia  
var. macradenia
click for image enlargement
©2009 Steve Matson

More photos of Eremogone macradenia var. macradenia
in CalPhotos

Geographic subdivisions for Eremogone macradenia var. macradenia:
s SN, SnGb, SnBr, SNE, DMoj
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map of distribution 1

(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurrence).


Data provided by the participants of the  Consortium of California Herbaria.

View all CCH records
All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.

CCH collections by month Flowering-Fruiting Monthly Counts

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).