Habit: Rhizome creeping and thin [to erect, trunk-like]
Leaf: pinnae deeply lobed, veins simple and free; resinous-glandular; winter-deciduous.
Etymology: (Greek: dark shield, in reference to the indusium)
Jepson eFlora Author: Susan Fawcett, John C. Game, Alan R. Smith & Thomas Lemieux
Reference: Fawcett & Smith 2021 A Generic Classification of the Thelypteridaceae. Fort Worth: Botanical Research Institute of Texas Press.
Unabridged Reference: Fawcett et al. 2021 Syst Bot 46:in press.Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)
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