Common Name: SUNFLOWER FAMILY Habit: Annual to tree. Leaf: basal and/or cauline, alternate, opposite, rarely whorled, simple to 2+ × compound. Inflorescence: 1° inflorescence a head, resembling a flower, of several types (see below), 1--many in generally +- cyme-like cluster; each head generally with +- calyx-like involucre of 1--many series of phyllaries (involucre bracts); receptacle of head flat to conic or columnar, paleate (bearing paleae = receptacle bracts) or epaleate; flowers 1--many per head. Flower: bisexual, unisexual, or sterile, +- small, of several types (see below); calyx 0 or modified into +- persistent pappus of bristles, scales, and/or awns; corolla radial or bilateral (0), lobes generally (0)3--5; stamens 4--5, filaments generally free, generally fused to corolla at tube/throat junction, anthers generally fused into cylinder around style, anther base generally rounded or cordate (deeply sagittate or with tail-like appendages), tip (= flattened appendage) generally projecting beyond pollen sac; pistil 1, 2-carpeled, ovary inferior, 1-chambered, 1-seeded, placenta basal, style 1, tip generally +- 2-branched (except in some staminate disk flowers), branch tips truncate or generally bearing +- brush-like appendages; stigmas 2, generally on adaxial faces of style branches. Fruit: achene (also called a cypsela) (drupe in Chrysanthemoides), cylindric to ovoid, sometimes compressed, generally deciduous with pappus attached. Genera In Family: +- 1500 genera, 23000 species: worldwide, many habitats. Note: Flower and head types differ in form and sexual condition. A disk flower has a generally radial corolla, with a cylindric tube, expanded throat, and generally 5 lobes. Disk flowers are generally bisexual and fertile but occasionally staminate with reduced ovaries. Discoid heads comprise only disk flowers. A radiant head is a variant of a discoid head, with peripheral disk flower corollas expanded, often bilateral. A ray flower corolla is bilateral, generally with a slender tube and flattened petal-like ray (single lip composed of generally 3 lobes). Ray flowers are generally pistillate or sterile (occasionally lacking styles). Radiate heads have peripheral ray flowers and central disk flowers. Disciform heads superficially resemble discoid heads, with pistillate or sterile flowers that lack rays, together with or separate from disk flowers. A ligulate flower is bisexual, with a bilateral, generally ephemeral corolla and 5-lobed ligule. Liguliflorous heads comprise only ligulate flowers. See glossary p. 31 for illustrations of family characteristics. Echinops sphaerocephalus L., Gaillardia aristata Pursh, Gaillardia pulchella Foug., Hymenothrix loomisii S.F. Blake, Tagetes erecta L., Thelesperma megapotamicum (Spreng.) Kuntze are waifs. Melampodium perfoliatum Kunth, historic urban waif. Ageratum conyzoides L., Guizotia abyssinica (L. f.) Cass., Santolina chamaecyparisus L., orth. var. are rare or uncommon escapes from cultivation. Dyssodia papposa, Ismelia carinata (Schousb.) Sch. Bip. [Chrysanthemum carinatum Schousb.], Mantisalca salmantica (L.) Briq. & Cavill. are historical or extirpated waifs in California. Inula helenium L. not documented in California. Taxa of Aster in TJM (1993) treated here in Almutaster, Doellingeria, Eurybia, Ionactis, Oreostemma, Sericocarpus, Symphyotrichum; Chamomilla in Matricaria; Bahia in Hymenothrix; Cnicus in Centaurea; Conyza in Erigeron and Laennecia; Dugaldia in Hymenoxys; Erechtites in Senecio; Hymenoclea in Ambrosia; Lembertia in Monolopia; Osteospermum ecklonis in Dimorphotheca; Picris echioides in Helminthotheca; Prionopsis in Grindelia; Raillardiopsis in Anisocarpus and Carlquistia; Schkuhria multiflora in Picradeniopsis; Trimorpha in Erigeron; Venidium in Arctotis; Viguiera in Aldama and Bahiopsis; Whitneya in Arnica. Amauriopsis in TJM2 (2012) treated here in Hymenothrix; Arida in Leucosyris; Bahia in Picradeniopsis; Eucephalus in Doellingeria. Unabridged Note: Largest family of vascular plants in California and of eudicots globally. eFlora Treatment Author: David J. Keil, except as noted Scientific Editor: David J. Keil, Bruce G. Baldwin.
Common Name: GROUNDSEL, RAGWORT, BUTTERWEED Habit: Biennial or perennial herb 3--100+ cm from rhizome or taproot with thin branched fibrous roots, loosely hairy to glabrous. Stem: 1--several. Leaf: pinnate or generally simple, basal and proximal cauline generally petioled, mid sessile, reduced, distal bract-like. Inflorescence: heads discoid or radiate, in compact or open cyme-like clusters, rarely single; involucre generally bell-shaped (cylindric); main phyllaries generally 8, 13, or 21 in 1 series, reflexed in fruit, green to +- red, linear, glabrous or hairy, subtended by a few reduced outer phyllaries, not black-tipped; receptacle epaleate. Ray Flower: 0--13(21); corolla pale yellow to deep orange-red. Disk Flower: 20--80+; corolla bell-shaped to tubular, lobes 5 erect to recurved, generally yellow (to deep orange-red); style tips truncate. Fruit: cylindric, generally prominently ribbed, glabrous or stiff-hairy; pappus of white minutely barbed bristles. Etymology: (J.G. Packer, Canadian botanist, b. 1929) Note: Formerly in Senecio. The common names groundsel, ragwort, and butterweed are applied to species of both Packera and Senecio. eFlora Treatment Author: Debra K. Trock Reference: Trock 2006 FNANM 20:570--602
Packera ionophylla (Greene) W.A. Weber & Á. Löve
NATIVE Habit: Perennial herb 1.5--3+ dm, from taprooted or branched creeping caudex. Stem: 1(2--3), cobwebby to woolly-tomentose or becoming glabrous. Leaf: basal and proximal cauline petioled, 1--3 cm, 1--2 cm wide, widely ovate or fiddle-shaped and pinnately lobed, lateral lobes 1--3 pairs, tapered to wedge-shaped at base, ultimate margins +- entire or crenate to coarsely dentate; distal cauline sessile, bract-like, entire. Inflorescence: heads radiate, 3--6; phyllaries (8)13 or 21, 7--10 mm, green, densely tomentose proximally, becoming glabrous distally, tips with tufted hairs. Ray Flower: 8--13; ray 8--10 mm. Disk Flower: 60--75+. Fruit: 2--2.5 mm, glabrous. Chromosomes: 2n=46. Ecology: Conifer woodland, dry, rocky slopes, granitic outcrops; Elevation: 1400--3000 m. Bioregional Distribution: s SN, SnGb, SnBr. Flowering Time: Jun--Aug Synonyms: Senecio ionophyllus Greene Jepson eFlora Author: Debra K. Trock Reference: Trock 2006 FNANM 20:570--602 Index of California Plant Names (ICPN; linked via the Jepson Online Interchange) Listed on CNPS Rare Plant Inventory Previous taxon: Packera indecora Next taxon: Packera layneae
Botanical illustration including Packera ionophylla
Citation for this treatment: Debra K. Trock 2012, Packera ionophylla, in Jepson Flora Project (eds.) Jepson eFlora, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=77396, accessed on December 03, 2024.
Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on December 03, 2024.
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Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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CCH collections by month
Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).