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Vascular Plants of California
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Holosteum umbellatum subsp. umbellatum

Higher Taxonomy
Family: CaryophyllaceaeView DescriptionDichotomous Key
Common Name: PINK FAMILY
Habit: Annual to perennial herb; rarely dioecious (Silene), taprooted or rhizome generally slender. Leaf: simple, generally opposite (subwhorled), entire, pairs at nodes often +- connected at bases; stipules generally 0; petiole generally 0. Inflorescence: generally cyme, generally open; flowers 1--many; involucre generally 0 (present in most Dianthus, Petrorhagia). Flower: generally bisexual, radial; hypanthium often present but obscure; sepals (4)5, +- free or fused into a tube, margins generally scarious, more so on inner 2 or not, tube generally not scarious, awns generally 0; petals (4)5 or 0, generally tapered to base (or with claw long, limb expanded), entire to 2--several-lobed, limb generally without scale-like appendages adaxially, generally without ear-like lobes at base; stamens generally 10, generally fertile, generally free, generally from ovary base; nectaries 0 or 5; ovary superior, generally 1-chambered, placentas basal or free-central, styles 2--5 with 0 branches or 1 with 2--3 branches. Fruit: capsule or utricle (rarely +- dehiscent), generally sessile. Seed: appendage generally 0 (present in Moehringia).
Genera In Family: +- 100 genera, 3000 species: widespread, especially arctic, alpine, temperate northern hemisphere; some cultivated (Agrostemma, Arenaria, Atocion, Cerastium, Dianthus, Gypsophila, Lychnis, Sagina, Saponaria, Silene). Note: Apetalous Caryophyllaceae can also be keyed in Rabeler & Hartman 2005 FNANM 5:5--8. Taxa of Minuartia in TJM2 treated here in Cherleria and Sabulina; Pseudostellaria in Hartmaniella and Torreyostellaria; Vaccaria in Gypsophila; Velezia in Dianthus.
eFlora Treatment Author: Ronald L. Hartman (deceased) & Richard K. Rabeler, except as noted
Scientific Editor: Bruce G. Baldwin & Thomas J. Rosatti.
Genus: HolosteumView Description 

Habit: Annual, ascending to erect, taprooted. Leaf: basal blades oblanceolate to spoon-shaped, short-petioled; cauline blades narrowly oblanceolate to oblong-ovate, sessile; vein 1. Inflorescence: terminal, umbel-like; flowers 3--12; involucre bracts 5--15, +- 0.3--1.8 mm; pedicels 0.5--2.5 cm. Flower: sepals 5, 2.5--4.5 mm, lanceolate to ovate, glabrous; petals 5, 3.3--5 mm, +- irregularly toothed; stamens 3--5; styles 3, 0.5--1.5 mm. Fruit: capsule, ovoid or ovoid-cylindric, straight; teeth 6, recurved. Seed: many, red-brown.
Etymology: (Greek: all bone, humorous reference to frailty of pl)
eFlora Treatment Author: Ronald L. Hartman (deceased) & Richard K. Rabeler
Reference: Rabeler & Hartman 2005 FNANM 5:94--95
Holosteum umbellatum L. subsp. umbellatum
Habit: Plant 5--25 cm. Stem: 1--many, unbranched, glandular-hairy +- near middle. Leaf: cauline 1--4 pairs in lower 1/2; blades 2--30 mm, glabrous or margin glandular-hairy. Inflorescence: bracts whorled, 1--2 mm, partly scarious; pedicels slender, reflexed in flower, erect in fruit. Flower: petals narrowly elliptic, white. Seed: 0.7--0.8 mm, compressed, with marginal ridge opposite elongate depression; tubercles low, rounded. Chromosomes: 2n=20,40.
Ecology: Uncommon. Disturbed areas, roadsides; Elevation: < 1270 m. Bioregional Distribution: KR, CaR, CCo?, MP; Distribution Outside California: to British Columbia, eastern North America; native to Europe; central and southwestern Asia; northern Africa. Flowering Time: Late winter--early spring
Unabridged Note: 4 other subspecies, native to Europe, central and southwestern Asia, northern Africa, introduced in South America, western Europe, southern Africa.
Jepson eFlora Author: Ronald L. Hartman (deceased) & Richard K. Rabeler
Reference: Rabeler & Hartman 2005 FNANM 5:94--95
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Botanical illustration including Holosteum umbellatum subsp. umbellatum

botanical illustration including Holosteum umbellatum subsp. umbellatum

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Citation for this treatment: Ronald L. Hartman (deceased) & Richard K. Rabeler 2012, Holosteum umbellatum subsp. umbellatum, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on February 21, 2024.

Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora,, accessed on February 21, 2024.

No expert verified images found for Holosteum umbellatum subsp. umbellatum.

Geographic subdivisions for Holosteum umbellatum subsp. umbellatum:
KR, CaR, CCo?, MP
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).


Data provided by the participants of the  Consortium of California Herbaria.
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All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.

CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).