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Vascular Plants of California
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Galium angustifolium subsp. foliosum


Higher Taxonomy
Family: RubiaceaeView DescriptionDichotomous Key
Common Name: MADDER FAMILY
Habit: Annual to tree, vine. Leaf: generally opposite (whorled), entire; stipules generally fused to stem, adjacent pairs occasionally fused, or occasionally leaf-like and appearing like whorled leaves. Inflorescence: cyme, panicle, spike, cluster, or flower 1, generally terminal and +- axillary. Flower: generally bisexual; calyx +- 4(5)-lobed, occasionally 0 (Galium, Crucianella) or 6 (Sherardia); corolla generally radial, 4(5)-lobed; stamens epipetalous, alternate corolla lobes, generally included; ovary generally inferior, chambers generally 2 or 4, style 1(2). Fruit: drupe, berry, or 2 or 4 nutlets [capsule].
Genera In Family: +- 500 genera, 6000 species: worldwide, especially tropics; many cultivated, including Coffea, coffee; Cinchona, quinine; many ornamental. Note: Diodia teres Walter doubtfully in California.
eFlora Treatment Author: Robert E. Preston & Lauramay T. Dempster, except as noted
Scientific Editor: Douglas H. Goldman, Bruce G. Baldwin.
Genus: GaliumView DescriptionDichotomous Key

Common Name: BEDSTRAW
Habit: Annual, perennial herb, occasionally subshrub, glabrous to hairy, generally scabrous; dioecious, bisexual, or flowers unisexual and bisexual. Stem: 4-angled, occasionally ridged lengthwise. Leaf: generally in whorls of >= 4, including leaf-like stipules. Inflorescence: panicles, axillary clusters (cymes), or occasionally 1 in axils. Flower: bisexual, or unisexual with sterile stamens or pistils; calyx 0; corolla generally rotate, occasionally +- bell-shaped, generally +- green, yellow to white, occasionally pink or red, lobes generally 4; ovary 2-lobed, styles 2, bases +- fused. Fruit: 2 nutlets or berry.
Etymology: (Greek: milk, from use of some species for curdling) Note: Ovary and fruit generally +- equally hairy on a pl; staminate plants generally identified by vestigial ovaries, pistillate plants generally by vestigial anthers. Galium saxatile L., Galium schultesii Vest, and Galium verum L. are lawn weeds in California.
eFlora Treatment Author: Valerie Soza
Reference: Dempster 1978 Univ Calif Publ Bot 73:1--33; Soza & Olmstead 2010 Amer J Bot 97:1630--1646
Unabridged Reference: Dempster 1973 Univ Calif Publ Bot 64:1--36; Dempster & Ehrendorfer 1965 Brittonia 17:289--334; Dempster & Stebbins 1968 Univ Calif Publ Bot 46:1--52; Dempster & Stebbins 1971 Madroño 21:71--95
Species: Galium angustifoliumView Description 


Common Name: NARROWLY LEAVED BEDSTRAW
Habit: Perennial herb, low, tufted or stems elongate, glabrous to hairy, at least base woody; dioecious. Stem: 6--100 cm. Leaf: in whorls of 4, +- strap-shaped, 3-veined. Inflorescence: panicle of few- to many-flowered clusters. Flower: corolla rotate, generally red to yellow. Fruit: nutlets, generally > pedicel; hairs dense, long, straight, spreading.

Galium angustifolium A. Gray subsp. foliosum (Hilend & J.T. Howell) Dempster & Stebbins
NATIVE
Habit: Plant +- woody above base, generally glabrous except leaf margin. Stem: 30--60 cm, ridges +- as wide as surfaces between; nodes short, obvious after leaf-fall. Leaf: generally 3--17 mm. Inflorescence: many-flowered, +- dense. Flower: corolla glabrous to puberulent. Chromosomes: 2n=22.
Ecology: Rocky slopes; Elevation: 30--60 m. Bioregional Distribution: n ChI. Flowering Time: Mar--Jul
Unabridged Synonyms: Galium angustifolium var. foliosum Hilend & Howell
Jepson eFlora Author: Valerie Soza
Reference: Dempster 1978 Univ Calif Publ Bot 73:1--33; Soza & Olmstead 2010 Amer J Bot 97:1630--1646
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)

Previous taxon: Galium angustifolium subsp. borregoense
Next taxon: Galium angustifolium subsp. gabrielense

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Citation for this treatment: Valerie Soza 2012, Galium angustifolium subsp. foliosum, in Jepson Flora Project (eds.) Jepson eFlora, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=50597, accessed on April 15, 2024.

Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on April 15, 2024.

Galium angustifolium subsp. foliosum
click for enlargement
©2006 Michael Charters
Galium angustifolium subsp. foliosum
click for enlargement
©2014 Steve Matson
Galium angustifolium subsp. foliosum
click for enlargement
©2006 Steve Matson
Galium angustifolium subsp. foliosum
click for enlargement
©2014 Steve Matson
Galium angustifolium subsp. foliosum
click for enlargement
©2006 Steve Matson

More photos of Galium angustifolium subsp. foliosum
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Geographic subdivisions for Galium angustifolium subsp. foliosum:
n ChI.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).





 

Data provided by the participants of the  Consortium of California Herbaria.
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View all CCH records
All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).