Common Name: BLADDERWORT FAMILY Habit: Annual, perennial herb, carnivorous, of moist or aquatic habitats. Stem: caudex or stolon, then often with thread-like branches. Leaf: simple, in rosette, or simple or dissected, emerging from caudex or stolon, with minute, carnivorous bladders +- throughout. Inflorescence: raceme or 1-flowered, scapose. Flower: bisexual; calyx lips 2[4], upper 3-lobed, lower 2-lobed, or lips unlobed; corolla 2-lipped, spurred at base, lower lip flat or arched upward, blocking throat or not; stamens 2, epipetalous; ovary superior, chamber 1, placenta generally free-central; stigma unequally 2-lobed, +- sessile. Fruit: capsule, round, 2-valved, circumscissile, or irregularly dehiscent. Seed: generally many, small. Genera In Family: 3 genera, 330 species: worldwide, especially tropics. eFlora Treatment Author: Barry A. Rice Scientific Editor: Thomas J. Rosatti.
Common Name: BLADDERWORT Habit: Carnivorous by bladders (here treated as modified leaves), into which small organisms are sucked when hairs at opening are triggered [epiphytic]. Stem: submersed or subterranean shoots [rarely caudex]; some aquatic species produce 2 kinds of stems, green (in water or at surface; leaves with flattened or thread-like segments; bladders 0--few) and white (generally buried in substrate; leaves 0; bladders many), the latter not always present in poor collections. Leaf: simple or generally dissected into narrow segments, alternate on stolon, margins often with bristles (visible at 10--30×). Inflorescence: raceme or 1-flowered, emergent; scape slender or wiry, bracts present. Flower: calyx lips 2[4], unlobed; corolla yellow [or not], with red-brown streaks or not, upper lip +- entire, lower entire or 3-lobed, spurred; rarely cleistogamous. Fruit: capsule. Etymology: (Latin: little bag, from bladders) Note: Size variable, often unreliable for identification; distinction between stems, leaves uncertain. Glands inside bladders consist of 2 pairs of oppositely directed arms, angles of divergence between which used (at 150× or more) to identify fresh or pressed specimens. Reference: Taylor 1989 Kew Bull Add Ser 14:1--724
Utricularia intermedia Hayne
NATIVE Stem: of 2 kinds, some freely floating, green, with leaves, rarely with bladders, others rooted in mud, white, without leaves, with bladders; winter buds bristly. Leaf: 3--15 mm, 3-parted at base, variously dissected above; ultimate segments < 20, +- linear, flattened, margins with 5--12(20) bristles, +- entire, tip obtuse. Inflorescence: 1--5-flowered; peduncle < 25 cm, < 1 mm diam; pedicel erect in fruit. Flower: corolla 10--16 mm; lower lip +- 2 × upper, +- > or +- = cylindric, acute spur. Fruit: circumscissile. Seed: not seen. Chromosomes: 2n=44. Ecology: Shallow (< 1 m) water; Elevation: 1200--2700 m. Bioregional Distribution: CaR, SNH, MP; Distribution Outside California: northern United States, Canada; circumboreal. Flowering Time: Jul--Sep Note: In some populations, divergence of bladder gland arm-pairs resembling that of Utricularia ochroleuca. Jepson eFlora Author: Barry A. Rice Reference: Taylor 1989 Kew Bull Add Ser 14:1--724 Index of California Plant Names (ICPN; linked via the Jepson Online Interchange) Listed on CNPS Rare Plant Inventory Previous taxon: Utricularia gibba Next taxon: Utricularia macrorhiza
Botanical illustration including Utricularia intermedia
Citation for this treatment: Barry A. Rice 2012, Utricularia intermedia, in Jepson Flora Project (eds.) Jepson eFlora, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=47604, accessed on April 19, 2024.
Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on April 19, 2024.
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(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).
Data provided by the participants of the
Consortium of California Herbaria.
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Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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CCH collections by month
Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).
