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Vascular Plants of California
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Pholistoma membranaceum


Higher Taxonomy
Family: HydrophyllaceaeView DescriptionDichotomous Key
Common Name: WATERLEAF FAMILY
Habit: Annual, perennial herb, generally hairy, generally taprooted. Stem: prostrate to erect. Leaf: simple to pinnately compound, basal or cauline, alternate or opposite; stipules 0. Inflorescence: cyme, generally raceme-like and coiled, or flowers 1. Flower: bisexual, generally radial; calyx lobes generally 5, generally fused at base, generally persistent, enlarging in fruit; corolla rotate to cylindric, generally deciduous, lobes generally 5, appendages in pairs on tube between filaments or 0; stamens generally 5, epipetalous, filament base sometimes appendaged, appendages scale-like; ovary generally superior, chambers 1 or 2, placentas 2, parietal, enlarged into chamber, sometimes meeting so ovary appears 2--5-chambered, style 1, style branches 2, stigmas generally head-like. Fruit: capsule, generally loculicidal; valves generally 2.
Genera In Family: 12 genera, 240--260 species: especially western US; some cultivated (Emmenanthe, Nemophila, Phacelia). Note: Included in Boraginaceae in TJM2 and some other treatments (Angiosperm Phylogeny Group IV 2016 Bot J Linn Soc 181:1--20) but treated as separate family (excluding Namaceae) by Boraginales Working Group (Luebert et al. 2016).
eFlora Treatment Author: Genevieve K. Walden, Robert W. Patterson & Richard R. Halse, except as specified
Scientific Editor: Bruce G. Baldwin
Genus: PholistomaView DescriptionDichotomous Key


Habit: Annual, fleshy. Stem: prostrate or reclined, many-branched, brittle; angles +- glabrous, bristly, or generally with hooked prickles. Leaf: cauline, lower opposite, upper alternate; petioles generally winged, clasping; blade pinnate-lobed, uppermost reduced, short-petioled, generally deltate, 3-lobed, with small, sharp bristles both surfaces. Inflorescence: terminal, axillary, opposite leaves, or flowers 1; pedicels present. Flower: calyx lobes hairy, bristly-ciliate; corolla rotate, lobes +- = tube, generally hairy; stamens included, equal, equally attached; ovary bristly-hairy, chamber appearing 1, style 1, 2-lobed in distal 1/2. Fruit: capsule, spheric, stout-bristly-hairy. Seed: 1--8, spheric, brown, pitted or honeycombed.
Species In Genus: 3 species: California, Arizona, Baja California. Etymology: (Greek: scale mouth)
eFlora Treatment Author: Robert W. Patterson & Richard R. Halse
Unabridged Reference: Constance 1939 Bull Torrey Bot Club 66:341--352
Pholistoma membranaceum (Benth.) Constance
NATIVE
Stem: 5--90 cm, generally glaucous. Leaf: petiole narrow-winged, not clasping; lower leaves 2--13 cm, 1--8 cm wide, blade oblong to ovate, base cordate or truncate, tip obtuse, lobes 5--11, oblong, obtuse, entire or 1-toothed. Inflorescence: flowers generally 2--10 in cymes; pedicel 5--20 mm. Flower: calyx lobes 1--3 mm, oblong, sinus appendages 0; corolla 3--6 mm, < 10 mm wide, white, lobe generally with purple spot; style 1--3 mm. Fruit: 2--4 mm wide, not enclosed in calyx. Seed: 1--2. Chromosomes: n=9.
Ecology: Beaches, bluffs, ravines, wooded slopes, desert washes; Elevation: 40--1400 m. Bioregional Distribution: c&s SNF, Teh, SnJV, CW, SW, D; Distribution Outside California: Baja California. Flowering Time: Feb--May
Jepson eFlora Author: Robert W. Patterson & Richard R. Halse
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)

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Botanical illustration including Pholistoma membranaceum

botanical illustration including Pholistoma membranaceum

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Citation for this treatment: Robert W. Patterson & Richard R. Halse 2021, Pholistoma membranaceum, in Jepson Flora Project (eds.) Jepson eFlora, Revision 9, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=37886, accessed on June 28, 2022.

Citation for the whole project: Jepson Flora Project (eds.) 2022, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on June 28, 2022.

Pholistoma membranaceum
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© 2012 Keir Morse
Pholistoma membranaceum
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© 2016 Barry Breckling
Pholistoma membranaceum
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© 2012 Keir Morse
Pholistoma membranaceum
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© 2014 Neal Kramer
Pholistoma membranaceum
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© 2014 Neal Kramer
Pholistoma membranaceum
click for enlargement
© 2013 Neal Kramer

More photos of Pholistoma membranaceum in CalPhotos



Geographic subdivisions for Pholistoma membranaceum:
c&s SNF, Teh, SnJV, CW, SW, D
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).





 

Data provided by the participants of the  Consortium of California Herbaria.
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Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).