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Pholistoma membranaceum


Higher Taxonomy
Family: HydrophyllaceaeView DescriptionDichotomous Key
Common Name: WATERLEAF FAMILY
Habit: Annual, perennial herb, generally hairy, generally taprooted. Stem: prostrate to erect. Leaf: simple to pinnately compound, basal or cauline, alternate or opposite; stipules 0. Inflorescence: cyme, generally raceme-like and coiled, or flowers 1. Flower: bisexual, generally radial; calyx lobes generally 5, generally fused at base, generally persistent, enlarging in fruit; corolla rotate to cylindric, generally deciduous, lobes generally 5, appendages in pairs on tube between filaments or 0; stamens generally 5, epipetalous, filament base sometimes appendaged, appendages scale-like; ovary generally superior, chambers 1 or 2, placentas 2, parietal, enlarged into chamber, sometimes meeting so ovary appears 2--5-chambered, style 1, style branches 2, stigmas generally head-like. Fruit: capsule, generally loculicidal; valves generally 2.
Genera In Family: 12 genera, 240--260 species: especially western US; some cultivated (Emmenanthe, Nemophila, Phacelia). Note: Included in Boraginaceae in TJM2 and some other treatments (Angiosperm Phylogeny Group IV 2016 Bot J Linn Soc 181:1--20) but treated as separate family (excluding Namaceae) by Boraginales Working Group (Luebert et al. 2016).
eFlora Treatment Author: Genevieve K. Walden, Robert W. Patterson & Richard R. Halse, except as specified
Scientific Editor: Bruce G. Baldwin
Genus: PholistomaView DescriptionDichotomous Key


Habit: Annual, fleshy. Stem: prostrate or reclined, many-branched, brittle; angles +- glabrous, bristly, or generally with hooked prickles. Leaf: cauline, lower opposite, upper alternate; petioles generally winged, clasping; blade pinnate-lobed, uppermost reduced, short-petioled, generally deltate, 3-lobed, with small, sharp bristles both surfaces. Inflorescence: terminal, axillary, opposite leaves, or flowers 1; pedicels present. Flower: calyx lobes hairy, bristly-ciliate; corolla rotate, lobes +- = tube, generally hairy; stamens included, equal, equally attached; ovary bristly-hairy, chamber appearing 1, style 1, 2-lobed in distal 1/2. Fruit: capsule, spheric, stout-bristly-hairy. Seed: 1--8, spheric, brown, pitted or honeycombed.
Etymology: (Greek: scale mouth)
eFlora Treatment Author: Robert W. Patterson & Richard R. Halse
Unabridged Reference: Constance 1939 Bull Torrey Bot Club 66:341--352
Pholistoma membranaceum (Benth.) Constance
NATIVE
Stem: 5--90 cm, generally glaucous. Leaf: petiole narrow-winged, not clasping; lower leaves 2--13 cm, 1--8 cm wide, blade oblong to ovate, base cordate or truncate, tip obtuse, lobes 5--11, oblong, obtuse, entire or 1-toothed. Inflorescence: flowers generally 2--10 in cymes; pedicel 5--20 mm. Flower: calyx lobes 1--3 mm, oblong, sinus appendages 0; corolla 3--6 mm, < 10 mm wide, white, lobe generally with purple spot; style 1--3 mm. Fruit: 2--4 mm wide, not enclosed in calyx. Seed: 1--2. Chromosomes: n=9.
Ecology: Beaches, bluffs, ravines, wooded slopes, desert washes; Elevation: 40--1400 m. Bioregional Distribution: c&s SNF, Teh, SnJV, CW, SW, D; Distribution Outside California: Baja California. Flowering Time: Feb--May
Jepson eFlora Author: Robert W. Patterson & Richard R. Halse
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)

Previous taxon: Pholistoma auritum var. auritum
Next taxon: Pholistoma racemosum

Botanical illustration including Pholistoma membranaceumbotanical illustration including Pholistoma membranaceum


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Citation for this treatment: Robert W. Patterson & Richard R. Halse 2021, Pholistoma membranaceum, in Jepson Flora Project (eds.) Jepson eFlora, Revision 9, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=37886, accessed on December 03, 2024.

Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on December 03, 2024.

Pholistoma membranaceum
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©2010 Thomas Stoughton
Pholistoma membranaceum
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©2016 Steve Matson
Pholistoma membranaceum
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©2008 Keir Morse
Pholistoma membranaceum
click for image enlargement
©2016 Steve Matson
Pholistoma membranaceum
click for image enlargement
©2008 Keir Morse

More photos of Pholistoma membranaceum
in CalPhotos



Geographic subdivisions for Pholistoma membranaceum:
c&s SNF, Teh, SnJV, CW, SW, D
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map of distribution 1

(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurrence).






 

Data provided by the participants of the  Consortium of California Herbaria.

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All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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CCH collections by month Flowering-Fruiting Monthly Counts

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).