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Vascular Plants of California
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Oreocarya flavoculata

Higher Taxonomy
Family: BoraginaceaeView DescriptionDichotomous Key
Habit: Annual, perennial herb, or shrub, often bristly or sharp-hairy. Stem: prostrate-decumbent to erect. Leaf: basal and/or cauline, simple, generally alternate, sometimes opposite, especially at base. Inflorescence: cymes, arranged singly or in groups of 2--5, generally coiled in flower, generally elongating in fruit. Flower: bisexual, generally radial; sepals 5, free or fused at least at base; corolla 5-lobed, salverform, funnel-shaped, rotate, or bell-shaped, appendages (often called "fornices") 0 or 5 at top of tube, when present often differentially pigmented, alternate stamens; stamens epipetalous; ovary superior, 4-lobed, style 1, entire or minutely 2-lobed (2-branched). Fruit: nutlets 1--4, when > 1, all similar (often called "homomorphic") or 1 or 2 dissimilar in size and/or shape from the others (often called "heteromorphic"), free (fused), smooth to roughened, prickly or bristly or not.
Genera In Family: +- 90 genera, +- 1600--1700 species: mostly temperate, especially western North America, Mediterranean; some cultivated (Borago, Echium, Myosotis, Symphytum). Toxicity: Many genera may be TOXIC from pyrrolizidine alkaloids or accumulated nitrates. Note: Sometimes still treated in broader sense of TJM2 (e.g., APG IV 2016 Bot J Linn Soc 181:1--20), but recent evidence (Luebert et al. 2016) supports segregation, for our flora, of the families Ehretiaceae, Heliotropiaceae, Hydrophyllaceae, Lennoaceae, and Namaceae.
eFlora Treatment Author: Michael G. Simpson, C. Matt Guilliams, Kristen Hasenstab-Lehman & Ronald B. Kelley
Scientific Editor: Bruce G. Baldwin, C. Matt Guilliams, Kristen Hasenstab-Lehman, David J. Keil, Ronald B. Kelley, Robert W. Patterson, Thomas J. Rosatti & Michael G. Simpson
Genus: OreocaryaView DescriptionDichotomous Key

Habit: Perennial or biennial herb, generally erect or decumbent. Stem: branches generally ascending to erect, hairy. Leaf: generally sessile; mostly in basal rosette, cauline alternate, distal reduced. Inflorescence: generally terminal, raceme- or panicle-like cymes, often densely clustered; bracts present or absent. Flower: generally persistent; sepals not fused; corolla tube 2--13 mm, limb 3--12 mm diam, white or yellow, appendages 5, yellow. Fruit: pedicel 0--12 mm in fruit; nutlets 1--4, lanceolate to wide-ovate, triangular, or elliptic, strongly bowed in profile in some, tip generally rounded, generally gray to brown, smooth, wrinkled (rugose), tubercled, or papillate, cross-ridged in some, with abaxial, longitudinal ridge present or not; margin rounded sharp-angled, or flat-rimmed; adaxially grooved above attachment scar, groove ending well below nutlet tip, edges gapped or abutted to +- overlapped, raised or not, generally forked or flared open at base; central fruit axis ("axis") extending to or beyond fruit.
Etymology: (Greek: mountain nut) Note: Generally homostylous, some spp. heterostylous. Segregated from Cryptantha based on molecular phylogenetic studies (Hasenstab-Lehman & Simpson 2012; Ripma et al. 2014; Simpson et al. 2017) and morphology, including perennial or biennial habit, relatively large nutlet size, and nutlet adaxial groove ending well below nutlet tip.
eFlora Treatment Author: Kristen E. Hasenstab-Lehman, Ronald B. Kelley, & Michael G. Simpson
Unabridged Reference: Johnston 1925 Contr Gray Herbarium 74:1--125; Payson 1927 Annals Missouri Bot. Gard. 14:211--358; Higgins 1971 Brigham Young Sci Bull Biol Ser 13:1--63, 1979 Great Basin Naturalist 39:293--350; Simpson & Hasenstab 2009 Crossosoma 35:1--59; Hasenstab-Lehman & Simpson 2012 Syst Bot 37:738--757; Ripma et al. 2014 Appl Plant Sci 2(12):1400062; Simpson et al. 2017 Taxon 66:1406--1420.
Oreocarya flavoculata A. Nelson
Habit: Perennial herb 5--35 cm, cespitose; caudex woody, branched. Stem: generally many, +- equal; strigose, bristly. Leaf: basal rosettes generally many; 2--11 cm; basal linear-oblanceolate to spoon-shaped, dense-silky-strigose, with bulbous-based bristles or not. Inflorescence: dense, cylindric or +- head-like, +- not elongated in fruit; pedicel not elongated in fruit, 1--1.5 mm. Flower: calyx 4--7 mm, 6--10 mm in fruit, with spreading, white or +- yellow bristles; corolla late-deciduous, tube 7--12 mm, limb 8--12 mm diam, appendages bright yellow. Fruit: nutlets 2--4, 2.5--3.2 mm, ovate to narrow-ovate, brown, dull, margin a +- flat narrow rim, +- arched in profile; abaxially dense-sharp-papillate, tubercled, cross-ridged, ridge present, rounded; adaxially tubercled, attachment scar edges gapped entire length, narrowed near middle, abruptly widened at base, edges raised; axis beyond nutlets.
Ecology: Common. Loose limestone-based soils; Elevation: 1200--3200 m. Bioregional Distribution: SNE, DMtns; Distribution Outside California: to Idaho, Colorado. Flowering Time: May--Jul Note: Heterostylous.
Synonyms: Cryptantha flavoculata (A. Nelson) Payson
Jepson eFlora Author: Kristen E. Hasenstab-Lehman, Ronald B. Kelley, & Michael G. Simpson
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)

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Citation for this treatment: Kristen E. Hasenstab-Lehman, Ronald B. Kelley, & Michael G. Simpson 2021, Oreocarya flavoculata, in Jepson Flora Project (eds.) Jepson eFlora, Revision 9,, accessed on June 16, 2024.

Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora,, accessed on June 16, 2024.

No expert verified images found for Oreocarya flavoculata.

Geographic subdivisions for Oreocarya flavoculata:
SNE, DMtns
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).


Data provided by the participants of the  Consortium of California Herbaria.
View all CCH records
All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.

CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).