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Vascular Plants of California
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Loeflingia squarrosa

Higher Taxonomy
Family: CaryophyllaceaeView DescriptionDichotomous Key
Common Name: PINK FAMILY
Habit: Annual to perennial herb; rarely dioecious (Silene), taprooted or rhizome generally slender. Leaf: simple, generally opposite (subwhorled), entire, pairs at nodes often +- connected at bases; stipules generally 0; petiole generally 0. Inflorescence: generally cyme, generally open; flowers 1--many; involucre generally 0 (present in most Dianthus, Petrorhagia). Flower: generally bisexual, radial; hypanthium often present but obscure; sepals (4)5, +- free or fused into a tube, margins generally scarious, more so on inner 2 or not, tube generally not scarious, awns generally 0; petals (4)5 or 0, generally tapered to base (or with claw long, limb expanded), entire to 2--several-lobed, limb generally without scale-like appendages adaxially, generally without ear-like lobes at base; stamens generally 10, generally fertile, generally free, generally from ovary base; nectaries 0 or 5; ovary superior, generally 1-chambered, placentas basal or free-central, styles 2--5 with 0 branches or 1 with 2--3 branches. Fruit: capsule or utricle (rarely +- dehiscent), generally sessile. Seed: appendage generally 0 (present in Moehringia).
Genera In Family: +- 100 genera, 3000 species: widespread, especially arctic, alpine, temperate northern hemisphere; some cultivated (Agrostemma, Arenaria, Atocion, Cerastium, Dianthus, Gypsophila, Lychnis, Sagina, Saponaria, Silene). Note: Apetalous Caryophyllaceae can also be keyed in Rabeler & Hartman 2005 FNANM 5:5--8. Taxa of Minuartia in TJM2 treated here in Cherleria and Sabulina; Pseudostellaria in Hartmaniella and Torreyostellaria; Vaccaria in Gypsophila; Velezia in Dianthus.
eFlora Treatment Author: Ronald L. Hartman (deceased) & Richard K. Rabeler, except as noted
Scientific Editor: Bruce G. Baldwin & Thomas J. Rosatti.
Genus: LoeflingiaView Description 

Habit: Annual, erect to prostrate, taprooted. Leaf: stipules 0.4--1.5 mm, bristle-like, scarious, entire, +- white; blade awl-like to oblong; vein 0--1. Inflorescence: axillary, sessile; flowers 1--2. Flower: sepals 5, 2.7--6 mm, +- free, +- lanceolate, glandular-hairy; petals 0 or vestigial; stamens 3--5; styles 3, < 0.1 mm. Fruit: capsule, 1.5--3.7 mm, lanceoloid to ovoid; valves 3, +- recurved at tip. Seed: many, tan with red-brown band on curved edge.
Etymology: (P. Loefling, Swedish botanist, explorer, 1729--1756)
eFlora Treatment Author: Ronald L. Hartman (deceased) & Richard K. Rabeler
Reference: Hartman & Rabeler 2005 FNANM 5:26--27
Loeflingia squarrosa Nutt.
Habit: Plant 1--12 cm, much-branched at base, glandular-hairy, +- fleshy. Stem: stiff. Leaf: blade 2--7 mm, erect to +- recurved, bristly; bases generally fused into a short scarious sheath; tip blunt to spine-tipped. Flower: cleistogamous; sepals awned, becoming hardened, margin often scarious, lateral spurs often present, bristly. Fruit: 0.5--0.8 × sepals, 3-angled. Seed: 0.4--0.7 mm, minutely papillate on flat edge.
Ecology: Sand, gravel of hills, mesas, dunes, disturbed areas; Elevation: < 1200 m. Bioregional Distribution: Teh, SnJV, SnFrB, SCo, PR, GB, DMoj; Distribution Outside California: to Oregon, Nebraska, Arkansas, Texas, northern Baja California. Flowering Time: Spring--summer
Synonyms: Loeflingia pusilla Curran; Loeflingia squarrosa subsp. artemisiarum Barneby & Twisselmann; Loeflingia squarrosa subsp. squarosa; Loeflingia squarrosa var. artemisiarum (Barneby & Twisselm.) Dorn Listed in CNPS Inventory; Loeflingia squarrosa var. squarrosa
Jepson eFlora Author: Ronald L. Hartman (deceased) & Richard K. Rabeler
Reference: Hartman & Rabeler 2005 FNANM 5:26--27
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)

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Botanical illustration including Loeflingia squarrosa

botanical illustration including Loeflingia squarrosa

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Citation for this treatment: Ronald L. Hartman (deceased) & Richard K. Rabeler 2012, Loeflingia squarrosa, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on June 25, 2024.

Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora,, accessed on June 25, 2024.

Loeflingia squarrosa
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©2014 Aaron Schusteff
Loeflingia squarrosa
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©2008 Keir Morse
Loeflingia squarrosa
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©2008 Keir Morse

More photos of Loeflingia squarrosa
in CalPhotos

Geographic subdivisions for Loeflingia squarrosa:
Teh, SnJV, SnFrB, SCo, PR, GB, DMoj
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).


Data provided by the participants of the  Consortium of California Herbaria.
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All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.

CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).