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Vascular Plants of California
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Hackelia velutina

Higher Taxonomy
Family: BoraginaceaeView DescriptionDichotomous Key
Habit: Annual to shrub or small tree, or non-green root parasite, often bristly or sharp-hairy. Stem: prostrate to erect. Leaf: basal and/or cauline, generally simple, generally alternate. Inflorescence: generally cymes, or panicle-, raceme-, head-, or spike-like, generally coiled in flower (often described as scorpioid), generally elongating in fruit, or flowers 1--2 per axil. Flower: bisexual, generally radial; sepals (4)5(10), fused at least at base, or free; corolla (4)5(10)-lobed, salverform, funnel-shaped, rotate, or bell-shaped, generally without scales at tube base, with 0 or 5 appendages at tube top, alternate stamens; stamens epipetalous; ovary generally superior, entire to 4-lobed, style 1(2), entire or 2-lobed or -branched. Fruit: valvate or circumscissile capsule or nutlets 1--4, free (fused), smooth to roughened, prickly or bristly or not.
Genera In Family: +- 120 genera, +- 2300 species: tropics, temperate, especially western North America, Mediterranean; some cultivated (Borago, Heliotropium, Echium, Myosotis, Nemophila, Phacelia, Symphytum, Wigandia). Toxicity: Many genera may be TOXIC from pyrrolizidine alkaloids or accumulated nitrates. Note: Recently treated to include Hydrophyllaceae, Lennoaceae. Wigandia urens added, as naturalized.
eFlora Treatment Author: Ronald B. Kelley, Robert Patterson, Richard R. Halse & Timothy C. Messick, family description, key to genera, treatment of genera by Ronald B. Kelley, except as noted
Scientific Editor: Ronald B. Kelley, Robert Patterson, Thomas J. Rosatti, Bruce G. Baldwin, David J. Keil.
Genus: HackeliaView DescriptionDichotomous Key

Common Name: STICKSEED
Habit: Perennial herb (biennial); hairs appressed to spreading; caudex generally branched in age, often +- woody, taprooted. Stem: ascending or erect. Leaf: lowest petioles generally +- = blades, +- winged, others 0. Inflorescence: coiled cymes, generally > 3, generally terminal and axillary, +- bracted; pedicel in fruit elongated, recurved to reflexed. Flower: calyx deep-5-lobed; corolla rotate-salverform, generally white with yellow patch adaxially, lobes appendaged near base. Fruit: nutlets erect, > style, attachment scar lateral-medial, generally with barb-tipped prickles abaxially and on margin.
Species In Genus: 40 species: generally western North America, southeastern Asia. Etymology: (J. Hackel, Czech botanist, 1783--1869) Note: Values for corolla limb diam take into account shrinkage during flower period. Difficult, study needed, especially in northern California, southeastern Asia; sometimes merged with Lappula.
Unabridged Note: In North America filling old-world role of flat-flowered forget-me-nots, Myosotis, Lepechiniella, etc, for pollinators.
eFlora Treatment Author: Ronald B. Kelley & Robert L. Carr
Unabridged Reference: Gentry & Carr 1976 Mem New York Bot Gard 26:121--227
Hackelia velutina (Piper) I.M. Johnst.
Stem: 4--8 dm; hairs generally spreading, +- dense below middle, +- dense to +- sparse above. Leaf: velvety-hairy; basal generally 5--17 cm, 0.5--2 cm wide, narrow-elliptic to oblanceolate; lower cauline reduced, ovate to narrow-lanceolate, +- clasping upward. Inflorescence: +- dense, branches several, few-flowered; pedicel 6--20 mm in fruit. Flower: calyx 2--3 mm; corolla tube generally >> calyx, generally +- purple, throat +- closed, limb 12--20 mm diam, generally blue to +- purple (pink), appendages long-exserted, much longer than wide, recurved, 2-pronged at tip, white; anthers hidden. Fruit: nutlets 4.5--6.5 mm, dull, roughened, abaxial prickles many, evenly distributed, +- = marginal.
Ecology: Dry, open slopes, forest clearings, roadsides; Elevation: 1350--2750 m. Bioregional Distribution: SNH; Distribution Outside California: western Nevada. Flowering Time: Jun--Aug
Synonyms: Hackelia longituba I.M. Johnst.
Unabridged Note: Though corolla distinctive, mistaken for Hackelia nervosa, Hackelia mundula: the unique long-exserted recurved corolla appendages and overall velvety hairs distinguish it from Hackelia nervosa; the exserted corolla tube, white corolla appendages, and dull, roughened nutlets separate it from Hackelia mundula. A CalPhoto image taken in southern SN shows a pink flowered pl, no doubt indicating introgression with Hackelia mundula. The suspected occurrence of Hackelia velutina in CaRH based on multiple CalPhoto images in the Lassen Volcanic National Park area, though it is uncertain whether voucher specimens exist.
Jepson eFlora Author: Ronald B. Kelley & Robert L. Carr
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)

Previous taxon: Hackelia sharsmithii
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Botanical illustration including Hackelia velutina

botanical illustration including Hackelia velutina


Citation for this treatment: Ronald B. Kelley & Robert L. Carr 2012, Hackelia velutina, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on May 10, 2021.

Citation for the whole project: Jepson Flora Project (eds.) 2021, Jepson eFlora,, accessed on May 10, 2021.

Hackelia velutina
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© 2009 Keir Morse
Hackelia velutina
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© 2009 Keir Morse
Hackelia velutina
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© 2015 Neal Kramer
Hackelia velutina
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© 2015 Neal Kramer
Hackelia velutina
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© 2015 Barry Rice
Hackelia velutina
click for enlargement
© 2009 Keir Morse

More photos of Hackelia velutina in CalPhotos

Geographic subdivisions for Hackelia velutina:
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).


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Data provided by the participants of the  Consortium of California Herbaria.
View all CCH records
All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.

CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).