Common Name: SUNFLOWER FAMILY Habit: Annual to tree. Leaf: basal and/or cauline, alternate, opposite, rarely whorled, simple to 2+ × compound. Inflorescence: 1° inflorescence a head, resembling a flower, of several types (see below), 1--many in generally +- cyme-like cluster; each head generally with +- calyx-like involucre of 1--many series of phyllaries (involucre bracts); receptacle of head flat to conic or columnar, paleate (bearing paleae = receptacle bracts) or epaleate; flowers 1--many per head. Flower: bisexual, unisexual, or sterile, +- small, of several types (see below); calyx 0 or modified into +- persistent pappus of bristles, scales, and/or awns; corolla radial or bilateral (0), lobes generally (0)3--5; stamens 4--5, filaments generally free, generally fused to corolla at tube/throat junction, anthers generally fused into cylinder around style, anther base generally rounded or cordate (deeply sagittate or with tail-like appendages), tip (= flattened appendage) generally projecting beyond pollen sac; pistil 1, 2-carpeled, ovary inferior, 1-chambered, 1-seeded, placenta basal, style 1, tip generally +- 2-branched (except in some staminate disk flowers), branch tips truncate or generally bearing +- brush-like appendages; stigmas 2, generally on adaxial faces of style branches. Fruit: achene (also called a cypsela) (drupe in Chrysanthemoides), cylindric to ovoid, sometimes compressed, generally deciduous with pappus attached. Genera In Family: +- 1500 genera, 23000 species: worldwide, many habitats. Note: Flower and head types differ in form and sexual condition. A disk flower has a generally radial corolla, with a cylindric tube, expanded throat, and generally 5 lobes. Disk flowers are generally bisexual and fertile but occasionally staminate with reduced ovaries. Discoid heads comprise only disk flowers. A radiant head is a variant of a discoid head, with peripheral disk flower corollas expanded, often bilateral. A ray flower corolla is bilateral, generally with a slender tube and flattened petal-like ray (single lip composed of generally 3 lobes). Ray flowers are generally pistillate or sterile (occasionally lacking styles). Radiate heads have peripheral ray flowers and central disk flowers. Disciform heads superficially resemble discoid heads, with pistillate or sterile flowers that lack rays, together with or separate from disk flowers. A ligulate flower is bisexual, with a bilateral, generally ephemeral corolla and 5-lobed ligule. Liguliflorous heads comprise only ligulate flowers. See glossary p. 31 for illustrations of family characteristics. Echinops sphaerocephalus L., Gaillardia aristata Pursh, Gaillardia pulchella Foug., Hymenothrix loomisii S.F. Blake, Tagetes erecta L., Thelesperma megapotamicum (Spreng.) Kuntze are waifs. Melampodium perfoliatum Kunth, historic urban waif. Ageratum conyzoides L., Guizotia abyssinica (L. f.) Cass., Santolina chamaecyparisus L., orth. var. are rare or uncommon escapes from cultivation. Dyssodia papposa, Ismelia carinata (Schousb.) Sch. Bip. [Chrysanthemum carinatum Schousb.], Mantisalca salmantica (L.) Briq. & Cavill. are historical or extirpated waifs in California. Inula helenium L. not documented in California. Taxa of Aster in TJM (1993) treated here in Almutaster, Doellingeria, Eurybia, Ionactis, Oreostemma, Sericocarpus, Symphyotrichum; Chamomilla in Matricaria; Bahia in Hymenothrix; Cnicus in Centaurea; Conyza in Erigeron and Laennecia; Dugaldia in Hymenoxys; Erechtites in Senecio; Hymenoclea in Ambrosia; Lembertia in Monolopia; Osteospermum ecklonis in Dimorphotheca; Picris echioides in Helminthotheca; Prionopsis in Grindelia; Raillardiopsis in Anisocarpus and Carlquistia; Schkuhria multiflora in Picradeniopsis; Trimorpha in Erigeron; Venidium in Arctotis; Viguiera in Aldama and Bahiopsis; Whitneya in Arnica. Amauriopsis in TJM2 (2012) treated here in Hymenothrix; Arida in Leucosyris; Bahia in Picradeniopsis; Eucephalus in Doellingeria. Unabridged Note: Largest family of vascular plants in California and of eudicots globally. eFlora Treatment Author: David J. Keil, except as noted Scientific Editor: David J. Keil, Bruce G. Baldwin.
Habit: [Subshrub], shrub. Stem: generally many from base. Leaf: alternate, generally drought-deciduous, simple, petioled, entire or rarely toothed. Inflorescence: heads radiate or discoid, 1 or in panicle-like cluster; peduncle generally long; involucre hemispheric; phyllaries graduated in 2--4+ series, free; receptacle paleate, palea folded around and falling with fruit. Ray Flower: sterile; style 0; ray yellow. Disk Flower: many; corolla yellow or brown-purple, tube slender, throat abruptly expanded, lobes triangular; anther tip ovate, +- acute; style tips triangular. Fruit: strongly compressed, obovate or wedge-shaped; edges long-ciliate; faces glabrous or short-hairy; pappus of 2 narrow scales or 0. Etymology: (Christoph Entzelt, German naturalist, 1517--1583) Note: Commonly hybridizing, especially in disturbed areas; Encelia farinosa × Encelia frutescens is common; Encelia californica × Encelia farinosa, Encelia actoni × Encelia farinosa, Encelia actoni × Encelia frutescens, Encelia frutescens × Encelia virginensis, Encelia farinosa × Geraea canescens have been reported. eFlora Treatment Author: David J. Keil & Curtis Clark Reference: Clark 2006 FNANM 21:118--122; Fehlberg & Ranker 2007 Syst Bot 32:692--699
Encelia actoni Elmer
NATIVE Habit: Shrub 5--15 dm, with many slender branches from base. Stem: branched below; young stem hairy; older stem with fissured bark. Leaf: scattered along stem; petiole 6--12 mm; blade 2.5--4 cm, ovate to deltate, acute, silvery green, canescent. Inflorescence: heads radiate, 1; peduncle canescent; involucre 8--14 mm; phyllaries ovate. Ray Flower: 14--25; ray 10--25 mm. Disk Flower: corolla 5--6 mm, yellow. Fruit: 5--7 mm; pappus generally 0. Chromosomes: 2n=36. Ecology: Open areas, rocky slopes, roadsides; Elevation: 200--2100 m. Bioregional Distribution: s SNF, s SnJV, n TR, e PR, W&I, s SNE, w D; Distribution Outside California: southwestern Nevada, northern Baja California. Flowering Time: Feb--Jul Synonyms: Encelia virginensis A. Nelson subsp. actoni (Elmer) D.D. Keck Jepson eFlora Author: David J. Keil & Curtis Clark Reference: Clark 2006 FNANM 21:118--122; Fehlberg & Ranker 2007 Syst Bot 32:692--699 Index of California Plant Names (ICPN; linked via the Jepson Online Interchange) Previous taxon: Encelia Next taxon: Encelia californica
Citation for this treatment: David J. Keil & Curtis Clark 2023, Encelia actoni, in Jepson Flora Project (eds.) Jepson eFlora, Revision 12, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=2562, accessed on December 03, 2024.
Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on December 03, 2024.
Geographic subdivisions for Encelia actoni:
s SNF, s SnJV, n TR, e PR, W&I, s SNE, w D
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(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurrence).
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Duplicates counted once; synonyms included.
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Blue line denotes eFlora flowering time (fruiting time in some monocot genera).