Common Name: WOOD FERN FAMILY Habit: Perennial herb, in soil or rock crevices; rhizome generally short-creeping, suberect, or erect, scales large, generally tan to brown, generally uniformly colored. Leaf: generally tufted, 5--200+ cm, generally +- alike; stipe generally firm, base generally darker, with many vascular strands; blade 1--4-pinnate, often with scales, hair-like scales, hairs (except clear, needle-like hairs generally 0), or short-stalked glands on axes, between veins or not, veins free to netted; rachis, costa generally grooved adaxially. Sporangia: sori round, along veins; indusia peltate or round-reniform; spores elliptic, winged, ridged, or spiny, scar linear. Genera In Family: +- 40--45 genera, > 1600 species: worldwide, especially tropics, wooded areas. Note: Based on molecular sequence data, Athyrium, Cystopteris, Woodsia removed to Woodsiaceae to preserve a monophyletic Dryopteridaceae. Unabridged Note: Current data (as reported by Schuettpelz & Pryer and in papers cited therein) suggest that Wooodsiaceae is paraphyletic (with respect to Aspeniaceae, Blechnaceae, and Thelypteridaceae), yet are insufficient to resolve the questions of circumscription (too few taxa, not enough genes sampled). Alternative classifications that would preserve monophyly include recognition of several additional, small families (e.g., Cystopteridaceae, Athyriaceae, and others not in California, each comprising just a few genera) or lumping at least 4 currently recognized families, many of long-standing use and acceptance; a conservative and expedient course is taken for now (Smith et al. 2006), pending further work. eFlora Treatment Author: Alan R. Smith Scientific Editor: Thomas J. Rosatti.
Common Name: WOOD FERN Habit: Rhizome short-creeping or ascending to suberect, stout. Leaf: stipe > 1.5 mm wide, firm, more densely scaly than midrib, base ×-section with many round vascular strands in an arc; blade >= 1--3-pinnate, proximal pinnae reduced or not, veins free, simple or forked; segments deeply pinnately lobed or not. Sporangia: sori round; indusium round-reniform, +- centrally attached at a sinus, generally persistent. Etymology: (Greek: oak, fern) Note: Hybrids unknown in California, frequent in eastern North America. Unabridged Reference: Montgomery & Paulton 1981 Fiddlehead Forum 8:25--31
Dryopteris arguta (Kaulf.) Watt
NATIVE Leaf: 30--60(100+) cm, 12--18(30) cm wide; stipe, midrib minutely glandular; blade lanceolate, 1--2-pinnate, proximal pinnae <= others, longest generally near blade base, sides +- equal, basiscopic pinnules 1--1.3 × acroscopic on same pinna, segments deeply pinnately lobed or not, teeth with bristle-like tips or not, veins into teeth; scales of pinna midribs lance-ovate to +- lanceolate. Chromosomes: 2n=82. Ecology: Locally common. Open, wooded slopes, caves; Elevation: < 2500 m. Bioregional Distribution: NW, SN, ScV (Sutter Buttes), CW, SW, MP (caves in Lava Beds National Monument), s DMoj; Distribution Outside California: to British Columbia, Arizona. Jepson eFlora Author: Alan R. Smith Index of California Plant Names (ICPN; linked via the Jepson Online Interchange) Previous taxon: Dryopteris Next taxon: Dryopteris expansa
Botanical illustration including Dryopteris arguta
Citation for this treatment: Alan R. Smith 2012, Dryopteris arguta, in Jepson Flora Project (eds.) Jepson eFlora, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=23524, accessed on December 02, 2024.
Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on December 02, 2024.
Geographic subdivisions for Dryopteris arguta:
NW, SN, ScV (Sutter Buttes), CW, SW, MP (caves in Lava Beds National Monument), s DMoj
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(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurrence).
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Duplicates counted once; synonyms included.
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