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Vascular Plants of California
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Brickellia knappiana


Higher Taxonomy
Family: Asteraceae (Compositae)View DescriptionDichotomous Key
Common Name: SUNFLOWER FAMILY
Habit: Annual to tree. Leaf: basal and/or cauline, alternate, opposite, rarely whorled, simple to 2+ × compound. Inflorescence: 1° inflorescence a head, resembling a flower, of several types (see below), 1--many in generally +- cyme-like cluster; each head generally with +- calyx-like involucre of 1--many series of phyllaries (involucre bracts); receptacle of head flat to conic or columnar, paleate (bearing paleae = receptacle bracts) or epaleate; flowers 1--many per head. Flower: bisexual, unisexual, or sterile, +- small, of several types (see below); calyx 0 or modified into +- persistent pappus of bristles, scales, and/or awns; corolla radial or bilateral (0), lobes generally (0)3--5; stamens 4--5, filaments generally free, generally fused to corolla at tube/throat junction, anthers generally fused into cylinder around style, anther base generally rounded or cordate (deeply sagittate or with tail-like appendages), tip (= flattened appendage) generally projecting beyond pollen sac; pistil 1, 2-carpeled, ovary inferior, 1-chambered, 1-seeded, placenta basal, style 1, tip generally +- 2-branched (except in some staminate disk flowers), branch tips truncate or generally bearing +- brush-like appendages; stigmas 2, generally on adaxial faces of style branches. Fruit: achene (also called a cypsela) (drupe in Chrysanthemoides), cylindric to ovoid, sometimes compressed, generally deciduous with pappus attached.
Genera In Family: +- 1500 genera, 23000 species: worldwide, many habitats. Note: Flower and head types differ in form and sexual condition. A disk flower has a generally radial corolla, with a cylindric tube, expanded throat, and generally 5 lobes. Disk flowers are generally bisexual and fertile but occasionally staminate with reduced ovaries. Discoid heads comprise only disk flowers. A radiant head is a variant of a discoid head, with peripheral disk flower corollas expanded, often bilateral. A ray flower corolla is bilateral, generally with a slender tube and flattened petal-like ray (single lip composed of generally 3 lobes). Ray flowers are generally pistillate or sterile (occasionally lacking styles). Radiate heads have peripheral ray flowers and central disk flowers. Disciform heads superficially resemble discoid heads, with pistillate or sterile flowers that lack rays, together with or separate from disk flowers. A ligulate flower is bisexual, with a bilateral, generally ephemeral corolla and 5-lobed ligule. Liguliflorous heads comprise only ligulate flowers. See glossary p. 31 for illustrations of family characteristics. Echinops sphaerocephalus L., Gaillardia aristata Pursh, Gaillardia pulchella Foug., Hymenothrix loomisii S.F. Blake, Tagetes erecta L., Thelesperma megapotamicum (Spreng.) Kuntze are waifs. Melampodium perfoliatum Kunth, historic urban waif. Ageratum conyzoides L., Guizotia abyssinica (L. f.) Cass., Santolina chamaecyparisus L., orth. var. are rare or uncommon escapes from cultivation. Dyssodia papposa, Ismelia carinata (Schousb.) Sch. Bip. [Chrysanthemum carinatum Schousb.], Mantisalca salmantica (L.) Briq. & Cavill. are historical or extirpated waifs in California. Inula helenium L. not documented in California. Taxa of Aster in TJM (1993) treated here in Almutaster, Doellingeria, Eurybia, Ionactis, Oreostemma, Sericocarpus, Symphyotrichum; Chamomilla in Matricaria; Bahia in Hymenothrix; Cnicus in Centaurea; Conyza in Erigeron and Laennecia; Dugaldia in Hymenoxys; Erechtites in Senecio; Hymenoclea in Ambrosia; Lembertia in Monolopia; Osteospermum ecklonis in Dimorphotheca; Picris echioides in Helminthotheca; Prionopsis in Grindelia; Raillardiopsis in Anisocarpus and Carlquistia; Schkuhria multiflora in Picradeniopsis; Trimorpha in Erigeron; Venidium in Arctotis; Viguiera in Aldama and Bahiopsis; Whitneya in Arnica. Amauriopsis in TJM2 (2012) treated here in Hymenothrix; Arida in Leucosyris; Bahia in Picradeniopsis; Eucephalus in Doellingeria.
Unabridged Note: Largest family of vascular plants in California and of eudicots globally.
eFlora Treatment Author: David J. Keil, except as noted
Scientific Editor: David J. Keil, Bruce G. Baldwin.
Genus: BrickelliaView DescriptionDichotomous Key


Common Name: BRICKELLBUSH
Habit: [Annual] perennial herb to shrub. Leaf: simple, alternate or opposite, generally resinous-dotted, veiny, main veins generally 3. Inflorescence: heads discoid, generally clustered; involucre cylindric to bell-shaped; phyllaries generally graduated, +- green, veiny-striate, spreading in age; receptacle generally flat, epaleate. Flower: corolla cylindric, +- white to pale yellow-green, occasionally tinged red or purple; anther tip ovate; style branches long, club-shaped, tips rounded. Fruit: 10-ribbed, generally cylindric, generally hairy; pappus of 10--40+ generally minutely barbed bristles, generally white.
Etymology: (John Brickell, early botanist in Georgia)
eFlora Treatment Author: Randall W. Scott
Reference: Scott 2006 FNANM 21:491--507
Unabridged Reference: King & Robinson 1987 Monogr Syst Bot Missouri Bot Gard 22:220--224
Brickellia knappiana Drew
NATIVE
Habit: Shrub 100--200 cm. Stem: arching, branched from near base, gland-dotted. Leaf: alternate, petiole 4--5 mm, blade 10--35 mm, 1--16 mm wide, lanceolate to lance-ovate, dentate to serrate, base acute to attenuate, tip acute to acuminate, faces densely gland-dotted and sparsely puberulent or hairy, not appearing glossy or varnished. Inflorescence: heads in panicle-like clusters, peduncle 2--5 mm, densely short-hairy and glandular, involucre 6--9 mm, cylindric to narrowly bell-shaped; phyllaries 18--22, in 5--7 series, 3--4-striate, tip acute to acuminate, margin scarious; outer lance-ovate, sparsely puberulent, gland-dotted, inner lanceolate, glabrous. Flower: 5--7; corolla 5--6.5 mm, white. Fruit: 2.5--3, puberulent to sparsely hairy; pappus bristles 28--32.
Ecology: Gravelly washes; Elevation: 700--1700 m. Bioregional Distribution: n&e DMtns (Panamint, n Kingston, Funeral, sw Argus ranges), DSon. Flowering Time: Sep--Oct Note: Possibly a hybrid between Brickellia californica and Brickellia longifolia var. multiflora or between Brickellia desertorum and Brickellia longifolia.
Unabridged Synonyms: Coleosanthus knappianus (Drew) Greene
Unabridged Note: Brickellia knappiana is noted as a possible hybrid between Brickellia californica and Brickellia multiflora (= Brickellia longifolia var. multiflora) or between Brickellia desertorum_ and Brickellia longifolia. Collections from mixed populations of Brickellia knappiana and Brickellia californica have been noted on herbarium specimens. Collections of specimens assignable to Brickellia knappiana that I have made have been where populations of Brickellia longifolia var. longifolia_and Brickellia desertorum intersect.
Jepson eFlora Author: Randall W. Scott
Reference: Scott 2006 FNANM 21:491--507
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)

Previous taxon: Brickellia incana
Next taxon: Brickellia longifolia

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botanical illustration including Brickellia knappiana

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Citation for this treatment: Randall W. Scott 2012, Brickellia knappiana, in Jepson Flora Project (eds.) Jepson eFlora, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=1815, accessed on April 19, 2024.

Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on April 19, 2024.

Brickellia knappiana
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©2013 Dana York
Brickellia knappiana
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©2013 Dana York
Brickellia knappiana
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©2013 Dana York
Brickellia knappiana
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©2016 Dana York
Brickellia knappiana
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©2016 Dana York

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Geographic subdivisions for Brickellia knappiana:
n&e DMtns (Panamint, n Kingston, Funeral, sw Argus ranges), DSon.
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).





 

Data provided by the participants of the  Consortium of California Herbaria.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).