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Vascular Plants of California
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Cardionema ramosissimum

Higher Taxonomy
Family: CaryophyllaceaeView DescriptionDichotomous Key
Common Name: PINK FAMILY
Habit: Annual to perennial herb; rarely dioecious (Silene), taprooted or rhizome generally slender. Leaf: simple, generally opposite (subwhorled), entire, pairs at nodes often +- connected at bases; stipules generally 0; petiole generally 0. Inflorescence: generally cyme, generally open; flowers 1--many; involucre generally 0 (present in Dianthus, Petrorhagia). Flower: generally bisexual, radial; hypanthium often present but obscure; sepals (4)5, +- free or fused into a tube, margins generally scarious, more so on inner 2 or not, tube generally not scarious, awns generally 0; petals (4)5 or 0, generally tapered to base (or with claw long, limb expanded), entire to 2--several-lobed, limb generally without scale-like appendages adaxially, generally without ear-like lobes at base; stamens generally 10, generally fertile, generally free, generally from ovary base; nectaries 0 or 5; ovary superior, generally 1-chambered, placentas basal or free-central, styles 2--5 with 0 branches or 1 with 2--3 branches. Fruit: capsule or utricle (rarely +- dehiscent), generally sessile. Seed: appendage generally 0 (present in Moehringia).
Genera In Family: 83 or 89 genera, 3000 species: widespread, especially arctic, alpine, temperate northern hemisphere; some cultivated (Agrostemma, Arenaria, Cerastium, Dianthus, Gypsophila, Lychnis, Sagina, Saponaria, Silene, Vaccaria). Note: Apetalous Caryophyllaceae can also be keyed in Rabeler & Hartman 2005 FNANM 5:5--8.
eFlora Treatment Author: Ronald L. Hartman & Richard K. Rabeler, except as noted
Scientific Editor: Thomas J. Rosatti.
Genus: CardionemaView Description 

Common Name: SANDMAT
Habit: Perennial herb, +- prostrate, taprooted. Leaf: stipules 4--8 mm, lanceolate to ovate, scarious, +- entire, white; blade needle-like; vein 1. Inflorescence: axillary; flowers 1--5, +- sessile. Flower: hypanthium cup-shaped, not abruptly expanded above; sepals 5, free, 1.2--2.8 mm (except awn), oblong to obovate, densely woolly, margin scarious below, awn 1.5--4 mm, stout, spine-tipped; petals 5, 0.3--0.5 mm, scale-like; stamens 3--5, on hypanthium rim; styles 2, 0.2 mm. Fruit: utricle, elliptic. Seed: 1, tan.
Species In Genus: 6 species: western North America, western South America. Etymology: (Greek: heart thread, from stamen shape)
Reference: Hartman 2005 FNANM 5:45--46
Cardionema ramosissimum (Weinm.) A. Nelson & J.F. Macbr.
Stem: 5--30+ cm, often +- covered by stipules. Leaf: 5--13 mm, finely spine-tipped, glabrous. Flower: sepal margin incurved near tip. Seed: 1.4--1.6 mm, narrowly ovate, not compressed.
Ecology: Sandy beaches, hills, dunes, bluffs; Elevation: < 400 m. Bioregional Distribution: NCo, CCo, SCo, n ChI, PR; Distribution Outside California: to Washington, Mexico; South America. Flowering Time: Spring--early summer
Jepson eFlora Author: Ronald L. Hartman & Richard K. Rabeler
Reference: Hartman 2005 FNANM 5:45--46
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)

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Botanical illustration including Cardionema ramosissimum

botanical illustration including Cardionema ramosissimum


Citation for this treatment: Ronald L. Hartman & Richard K. Rabeler 2012, Cardionema ramosissimum, in Jepson Flora Project (eds.) Jepson eFlora, /eflora/eflora_display.php?tid=17198, accessed on August 10, 2020.

Citation for the whole project: Jepson Flora Project (eds.) 2020, Jepson eFlora,, accessed on August 10, 2020.

Cardionema ramosissimum
click for enlargement
© 2006 Steve Matson
Cardionema ramosissimum
click for enlargement
© 2006 Steve Matson
Cardionema ramosissimum
click for enlargement
© 2018 Neal Kramer
Cardionema ramosissimum
click for enlargement
© 2006 Steve Matson
Cardionema ramosissimum
click for enlargement
© 2011 Neal Kramer
Cardionema ramosissimum
click for enlargement
© 2015 Barry Breckling

More photos of Cardionema ramosissimum in CalPhotos

Geographic subdivisions for Cardionema ramosissimum:
NCo, CCo, SCo, n ChI, PR
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).


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Data provided by the participants of the  Consortium of California Herbaria.
View all CCH records
All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.

CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).