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Vascular Plants of California
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Berula erecta

Higher Taxonomy
Family: Apiaceae (Umbelliferae)View DescriptionDichotomous Key
Habit: Annual to perennial herb [shrub, tree], generally from taproot. Stem: generally +- scapose, generally ribbed, hollow. Leaf: basal and generally cauline, generally alternate; stipules generally 0; petiole base generally sheathing stem; blade generally much dissected, occasionally compound. Inflorescence: umbel or head, simple or compound, generally peduncled; bracts present in involucres or 0; bractlets generally present in "involucels". Flower: many, small, generally bisexual (or some staminate), generally radial (or outer bilateral); calyx 0 or lobes 5, small; petals 5, free, generally ovate or spoon-shaped, generally incurved at tips, generally +- ephemeral; stamens 5; pistil 1, ovary inferior, 2-chambered, generally with a +- conic, persistent projection or platform at tip subtending 2 free styles. Fruit: 2 dry, 1-seeded halves (= mericarps), separating from each other but generally +- persistent to central axis; ribs on halves 5, 2 marginal, 3 to back; oil tubes 1--several per interval between ribs.
Genera In Family: 300 genera, 3000 species: +- worldwide, especially temperate; many cultivated for food or spice (e.g., Carum, caraway; Daucus; Petroselinum); Bupleurum lancifolium Hornem. is historical garden weed; some toxic (e.g., Conium). Note: Mature fruit generally critical in identification, shape given in outline. Hydrocotyle moved to Araliaceae, Orogenia moved to Lomatium, Sphenosciadium moved to Angelica. Petroselinum crispum (Mill.) A.W. Hill is a waif.
eFlora Treatment Author: Lincoln Constance & Margriet Wetherwax, except as noted
Scientific Editor: Douglas H. Goldman, Bruce G. Baldwin.
Genus: BerulaView Description 


Species In Genus: 1 sp. Etymology: (Latin: water-cress)
eFlora Treatment Author: Lincoln Constance & Margriet Wetherwax
Berula erecta (Huds.) Coville
Habit: Perennial herb, < 15 dm, stoloned, glabrous. Stem: ascending or erect, branched, hollow, rooting at proximal nodes. Leaf: petiole 4--12 cm, narrowly sheathing; blade 1.5 dm, oblong, 1-pinnate, much dissected if submerged, leaflets 7--12 pairs, 1--8 cm, oblong to ovate, sessile, lobed or serrate. Inflorescence: umbels compound, peduncled, terminal or opposite leaves; bracts, bractlets leaf-like, lanceolate; rays, pedicels +- unequal, spreading. Flower: calyx lobes minute, persistent; petals wide, white. Fruit: 1.5--2 mm, +- round, compressed side-to-side, glabrous; ribs thread-like, inconspicuous in corky fruit wall; oil tubes apparent, deeply embedded in fruit wall; fruit axis divided to base, branches adhering to and falling with fruit-halves. Chromosomes: 2n=18.
Ecology: Marshy areas, streams; Elevation: < 2000 m. Bioregional Distribution: CA; Distribution Outside California: to British Columbia, New York, Texas, Mexico; Eurasia, Africa. Toxicity: Possibly TOXIC: possibly poisonous to livestock. Flowering Time: Jul--Oct
Jepson eFlora Author: Lincoln Constance & Margriet Wetherwax
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)

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Botanical illustration including Berula erecta

botanical illustration including Berula erecta


Citation for this treatment: Lincoln Constance & Margriet Wetherwax 2012, Berula erecta, in Jepson Flora Project (eds.) Jepson eFlora,, accessed on April 11, 2021.

Citation for the whole project: Jepson Flora Project (eds.) 2021, Jepson eFlora,, accessed on April 11, 2021.

Berula erecta
click for enlargement
© 2012 Steve Matson
Berula erecta
click for enlargement
© 2016 Steve Matson
Berula erecta
click for enlargement
© 2015 Steve Matson
Berula erecta
click for enlargement
© 2012 Steve Matson
Berula erecta
click for enlargement
© 2012 Steve Matson
Berula erecta
click for enlargement
© 2015 Steve Matson

More photos of Berula erecta in CalPhotos

Geographic subdivisions for Berula erecta:
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).


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Data provided by the participants of the  Consortium of California Herbaria.
View all CCH records
All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.

CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).