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Vascular Plants of California
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Johnstonella racemosa
SHRUBBY JOHNSTONELLA


Higher Taxonomy
Family: BoraginaceaeView DescriptionDichotomous Key
Common Name: BORAGE FAMILY
Habit: Annual, perennial herb, or shrub, often bristly or sharp-hairy. Stem: prostrate-decumbent to erect. Leaf: basal and/or cauline, simple, generally alternate, sometimes opposite, especially at base. Inflorescence: cymes, arranged singly or in groups of 2--5, generally coiled in flower, generally elongating in fruit. Flower: bisexual, generally radial; sepals 5, free or fused at least at base; corolla 5-lobed, salverform, funnel-shaped, rotate, or bell-shaped, appendages (often called "fornices") 0 or 5 at top of tube, when present often differentially pigmented, alternate stamens; stamens epipetalous; ovary superior, 4-lobed, style 1, entire or minutely 2-lobed (2-branched). Fruit: nutlets 1--4, when > 1, all similar (often called "homomorphic") or 1 or 2 dissimilar in size and/or shape from the others (often called "heteromorphic"), free (fused), smooth to roughened, prickly or bristly or not.
Genera In Family: +- 90 genera, +- 1600--1700 species: mostly temperate, especially western North America, Mediterranean; some cultivated (Borago, Echium, Myosotis, Symphytum). Toxicity: Many genera may be TOXIC from pyrrolizidine alkaloids or accumulated nitrates. Note: Sometimes still treated in broader sense of TJM2 (e.g., APG IV 2016 Bot J Linn Soc 181:1--20), but recent evidence (Luebert et al. 2016) supports segregation, for our flora, of the families Ehretiaceae, Heliotropiaceae, Hydrophyllaceae, Lennoaceae, and Namaceae.
eFlora Treatment Author: Michael G. Simpson, C. Matt Guilliams, Kristen Hasenstab-Lehman & Ronald B. Kelley
Scientific Editor: Bruce G. Baldwin, C. Matt Guilliams, Kristen Hasenstab-Lehman, David J. Keil, Ronald B. Kelley, Robert W. Patterson, Thomas J. Rosatti & Michael G. Simpson
Genus: JohnstonellaView DescriptionDichotomous Key


Habit: Annual, sometimes appearing perennial. Stem: branches erect to decumbent, hairy, sometimes stout and woody at base. Leaf: cauline only, alternate, sessile, linear to elliptic, hairy. Inflorescence: terminal, of raceme- or spike-like cymes, in groups of 1--2, basal bracts in some. Flower: bracts present or 0; sepals free to base; corolla limb 0.5--4 mm diam, white, appendages generally 5, yellow. Fruit: pedicel 0--4(--6) mm in fruit; nutlets (3)4, similar or dissimilar, if dissimilar single different nutlet positioned away from inflorescence axis, nutlets smooth to white-tubercled, margin rounded, sharp-edged, or a +- flat rim or wing; abaxial ridge 0; adaxially grooved above attachment scar, scar generally lateral, narrow, open to closed, generally forked or flared open at base; style generally extended beyond nutlets.
Species In Genus: +- 16 species: western North America, western and southern South America. Etymology: (Ivan M. Johnston, American botanist, 1898--1960) Note: A segregate of Cryptantha, Johnstonella is strongly supported as a separate lineage by molecular phylogenetic studies (Hasenstab-Lehman & Simpson 2012; Simpson et al. 2017).
eFlora Treatment Author: Michael G. Simpson, Kristen E. Hasenstab-Lehman & Ronald B. Kelley
Reference: Johnston 1925 Contr Gray Herbarium 74:1--125; Simpson & Hasenstab 2009 Crossosoma 35:1--59; Hasenstab-Lehman & Simpson 2012 Syst Bot 37:738--757; Simpson et al. 2014 Taxon 63:930--931; Simpson et al. 2017 Taxon 66:1406--1420; Simpson et al. 2019 Phytotaxa 425:279--289; Simpson et al. 2020 Madroño 67:5--8.
Johnstonella racemosa (S. Watson ex A. Gray) Brand
NATIVE
Habit: Annual, often woody at base and resembling perennial, 20--100 cm, generally flowering 1st year, becoming rounded, shrub-like. Stem: branches many, throughout, spreading to ascending, those distal +- wiry; canescent and sparse-spreading-bristly. Leaf: 1.5--4(--6) cm, narrow-oblanceolate to linear, bulbous-based-bristly. Inflorescence: raceme-like, generally in 1s, open, few-flowered; bracts minute; pedicel elongated in fruit, 1--4(--6) mm, +- loosely spreading. Flower: calyx 1--2 mm, 2--4 mm in fruit, +- persistent, lobes lance-linear, bristly in lower 1/2, bristles spreading to reflexed, midvein thickened; corolla persistent, limb 1--3 mm diam, flat, appendages yellow. Fruit: nutlets 3--4, 0.8--2 mm, 1 > others, persistent, long-ovate, dark gray with pale tubercles, +- shiny, margin a +- flat rim, wider near tip; abaxially low-rounded, sparsely white-fine-tubercled, ridge 0; adaxially biconvex, sparse-tubercled, attachment scar edges not raised, +- abutted near tip, wide-triangular-gapped.
Ecology: Rocky slopes, crevices, washes, canyons, desert scrub; Elevation: < 1400(1670) m. Bioregional Distribution: n SnBr, e PR, n W&I, D; Distribution Outside California: southern Nevada, western Arizona, Baja California. Flowering Time: Feb--May
Synonyms: Cryptantha racemosa (A. Gray) Greene
Jepson eFlora Author: Michael G. Simpson, Kristen E. Hasenstab-Lehman & Ronald B. Kelley
Reference: Johnston 1925 Contr Gray Herbarium 74:1--125; Simpson & Hasenstab 2009 Crossosoma 35:1--59; Hasenstab-Lehman & Simpson 2012 Syst Bot 37:738--757; Simpson et al. 2014 Taxon 63:930--931; Simpson et al. 2017 Taxon 66:1406--1420; Simpson et al. 2019 Phytotaxa 425:279--289; Simpson et al. 2020 Madroño 67:5--8.
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Citation for this treatment: Michael G. Simpson, Kristen E. Hasenstab-Lehman & Ronald B. Kelley 2021, Johnstonella racemosa, in Jepson Flora Project (eds.) Jepson eFlora, Revision 9, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=101946, accessed on May 19, 2022.

Citation for the whole project: Jepson Flora Project (eds.) 2022, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on May 19, 2022.

No expert verified images found for Johnstonella racemosa.



Geographic subdivisions for Johnstonella racemosa:
n SnBr, e PR, n W&I, D
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).





 

Data provided by the participants of the  Consortium of California Herbaria.
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All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).