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Greeneocharis circumscissa var. rosulata
ROSETTE CUSHION GREENEOCHARIS


Higher Taxonomy
Family: BoraginaceaeView DescriptionDichotomous Key
Common Name: BORAGE FAMILY
Habit: Annual, perennial herb, or shrub, often bristly or sharp-hairy. Stem: prostrate-decumbent to erect. Leaf: basal and/or cauline, simple, generally alternate, sometimes opposite, especially at base. Inflorescence: cymes, arranged singly or in groups of 2--5, generally coiled in flower, generally elongating in fruit. Flower: bisexual, generally radial; sepals 5, free or fused at least at base; corolla 5-lobed, salverform, funnel-shaped, rotate, or bell-shaped, appendages (often called "fornices") 0 or 5 at top of tube, when present often differentially pigmented, alternate stamens; stamens epipetalous; ovary superior, 4-lobed, style 1, entire or minutely 2-lobed (2-branched). Fruit: nutlets 1--4, when > 1, all similar (often called "homomorphic") or 1 or 2 dissimilar in size and/or shape from the others (often called "heteromorphic"), free (fused), smooth to roughened, prickly or bristly or not.
Genera In Family: +- 90 genera, +- 1600--1700 species: mostly temperate, especially western North America, Mediterranean; some cultivated (Borago, Echium, Myosotis, Symphytum). Toxicity: Many genera may be TOXIC from pyrrolizidine alkaloids or accumulated nitrates. Note: Sometimes still treated in broader sense of TJM2 (e.g., APG IV 2016 Bot J Linn Soc 181:1--20), but recent evidence (Luebert et al. 2016) supports segregation, for our flora, of the families Ehretiaceae, Heliotropiaceae, Hydrophyllaceae, Lennoaceae, and Namaceae.
eFlora Treatment Author: Michael G. Simpson, C. Matt Guilliams, Kristen Hasenstab-Lehman & Ronald B. Kelley
Scientific Editor: Bruce G. Baldwin, C. Matt Guilliams, Kristen Hasenstab-Lehman, David J. Keil, Ronald B. Kelley, Robert W. Patterson, Thomas J. Rosatti & Michael G. Simpson
Genus: GreeneocharisView DescriptionDichotomous Key

Habit: Annual herb, cushion-like, roots, plant bases red-purple or not. Stem: branches ascending to erect, slender, generally strigose. Leaf: sessile, alternate, congested at branch tips, linear, oblanceolate, or narrowly oblong, hairy. Inflorescence: spike-like cymes; flower bracts present. Flower: calyx lobes fused at base, tube circumscissile in fruit; corolla limb 1--6 mm diam, white, appendages present. Fruit: pedicel 0--0.5 mm in fruit; fruit axis +- 1/2--1× nutlet length; nutlets (2)3 or 4, generally similar (dissimilar), smooth to roughened, ridge 0; margin rounded; attachment scar abutted near apex, forked and gapped at base; style extended to or just below nutlet tips.
Species In Genus: 2 species: western North America. Etymology: (Greek: Greene's delight, for Edward L. Greene, California botanist, 1843--1915) Note: Greeneocharis is a segregate of Cryptantha, strongly supported as a separate lineage by molecular phylogenetic studies.
eFlora Treatment Author: Ronald B. Kelley, Kristen Hasenstab-Lehman, & Michael G. Simpson
Reference: Johnston 1925 Contr Gray Herbarium 74:1--125; Mathew & Raven 1962 Madroño 16:168--171; Simpson & Hasenstab 2009 Crossosoma 35:1--59; Hasenstab-Lehman & Simpson 2012 Syst Bot 37: 738--757; Simpson et al. 2017 Taxon 66:1406--1420
Species: Greeneocharis circumscissaView Description 


Habit: Annual 0.5--10 cm diam, rounded to cushion-like. Stem: 0.5--8 cm, branches slender; strigose and bristly or rough-hairy, hairs generally ascending to occasionally spreading. Leaf: 0.3--1.5 cm; hairs rough, generally ascending to spreading. Inflorescence: in 1s to 3s, 1--5-flowered, dense in fruit; bracts present; pedicel +- 0.5 mm, erect. Flower: calyx circumscissile below middle, cup-like in fruit, hairs +- like leaf hairs; corolla limb generally 1--2 mm diam, appendages inconspicuous, white to pale yellow. Fruit: nutlets +- flattened, base truncate; abaxial ridge 0; adaxial attachment scar edges not raised, abutted near tip.

Greeneocharis circumscissa (Hook. & Arn.) Rydb. var. rosulata (J.T. Howell) Hasenstab & M.G. Simpson
NATIVE
Habit: Plant 0.5--3.5 cm, generally prostrate; root, plant base not red-purple. Stem: 0.5--2 cm, branches few, generally compact; strigose, hairs generally ascending. Leaf: oblanceolate to narrow-oblong, margin +- flat. Flower: calyx 2.5--3 mm in fruit; corolla appendages minute, white. Fruit: nutlets (2)3 or 4, 1.5--1.8 mm, similar, lance-triangular, smooth, shiny, gray, margin a +- flat linear rim, tip acuminate; abaxially +- flat; adaxially biconvex, attachment scar edges occasionally gapped at base; axis not to nutlet tips.
Ecology: Barren granitic gravels; Elevation: 2950--3650 m. Bioregional Distribution: SNH (Inyo, Tulare cos.). Flowering Time: Jul--Aug
Synonyms: Cryptantha circumscissa (Hook. & Arn.) I.M. Johnst. var. rosulata J.T. Howell
Jepson eFlora Author: Ronald B. Kelley, Kristen Hasenstab-Lehman, & Michael G. Simpson
Reference: Johnston 1925 Contr Gray Herbarium 74:1--125; Mathew & Raven 1962 Madroño 16:168--171; Simpson & Hasenstab 2009 Crossosoma 35:1--59; Hasenstab-Lehman & Simpson 2012 Syst Bot 37: 738--757; Simpson et al. 2017 Taxon 66:1406--1420
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)
Listed on CNPS Rare Plant Inventory

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Citation for this treatment: Ronald B. Kelley, Kristen Hasenstab-Lehman, & Michael G. Simpson 2021, Greeneocharis circumscissa var. rosulata, in Jepson Flora Project (eds.) Jepson eFlora, Revision 9, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=101725, accessed on May 19, 2022.

Citation for the whole project: Jepson Flora Project (eds.) 2022, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on May 19, 2022.

No expert verified images found for Greeneocharis circumscissa var. rosulata.



Geographic subdivisions for Greeneocharis circumscissa var. rosulata:
SNH (Inyo, Tulare cos.).
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).





 

Data provided by the participants of the  Consortium of California Herbaria.
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Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).