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Greeneocharis circumscissa var. rosulata

Higher Taxonomy
Family: BoraginaceaeView DescriptionDichotomous Key
Habit: Annual, perennial herb, or shrub, often bristly or sharp-hairy. Stem: prostrate-decumbent to erect. Leaf: basal and/or cauline, simple, generally alternate, sometimes opposite, especially at base. Inflorescence: cymes, arranged singly or in groups of 2--5, generally coiled in flower, generally elongating in fruit. Flower: bisexual, generally radial; sepals 5, free or fused at least at base; corolla 5-lobed, salverform, funnel-shaped, rotate, or bell-shaped, appendages (often called "fornices") 0 or 5 at top of tube, when present often differentially pigmented, alternate stamens; stamens epipetalous; ovary superior, 4-lobed, style 1, entire or minutely 2-lobed (2-branched). Fruit: nutlets 1--4, when > 1, all similar (often called "homomorphic") or 1 or 2 dissimilar in size and/or shape from the others (often called "heteromorphic"), free (fused), smooth to roughened, prickly or bristly or not.
Genera In Family: +- 90 genera, +- 1600--1700 species: mostly temperate, especially western North America, Mediterranean; some cultivated (Borago, Echium, Myosotis, Symphytum). Toxicity: Many genera may be TOXIC from pyrrolizidine alkaloids or accumulated nitrates. Note: Sometimes still treated in broader sense of TJM2 (e.g., APG IV 2016 Bot J Linn Soc 181:1--20), but recent evidence (Luebert et al. 2016) supports segregation, for our flora, of the families Ehretiaceae, Heliotropiaceae, Hydrophyllaceae, Lennoaceae, and Namaceae.
eFlora Treatment Author: Michael G. Simpson, C. Matt Guilliams, Kristen Hasenstab-Lehman & Ronald B. Kelley
Scientific Editor: Bruce G. Baldwin, C. Matt Guilliams, Kristen Hasenstab-Lehman, David J. Keil, Ronald B. Kelley, Robert W. Patterson, Thomas J. Rosatti & Michael G. Simpson
Genus: GreeneocharisView DescriptionDichotomous Key

Habit: Annual herb, cushion-like, roots, plant bases red-purple or not. Stem: branches ascending to erect, slender, generally strigose. Leaf: sessile, alternate, congested at branch tips, linear, oblanceolate, or narrowly oblong, hairy. Inflorescence: spike-like cymes; flower bracts present. Flower: calyx lobes fused at base, tube circumscissile in fruit; corolla limb 1--6 mm diam, white, appendages present. Fruit: pedicel 0--0.5 mm in fruit; fruit axis +- 1/2--1× nutlet length; nutlets (2)3 or 4, generally similar (dissimilar), smooth to roughened, ridge 0; margin rounded; attachment scar abutted near apex, forked and gapped at base; style extended to or just below nutlet tips.
Etymology: (Greek: Greene's delight, for Edward L. Greene, California botanist, 1843--1915) Note: Greeneocharis is a segregate of Cryptantha, strongly supported as a separate lineage by molecular phylogenetic studies.
eFlora Treatment Author: Ronald B. Kelley, Kristen Hasenstab-Lehman, & Michael G. Simpson
Reference: Johnston 1925 Contr Gray Herbarium 74:1--125; Mathew & Raven 1962 Madroño 16:168--171; Simpson & Hasenstab 2009 Crossosoma 35:1--59; Hasenstab-Lehman & Simpson 2012 Syst Bot 37: 738--757; Simpson et al. 2017 Taxon 66:1406--1420
Species: Greeneocharis circumscissaView Description 

Habit: Annual 0.5--10 cm diam, rounded to cushion-like. Stem: 0.5--8 cm, branches slender; strigose and bristly or rough-hairy, hairs generally ascending to occasionally spreading. Leaf: 0.3--1.5 cm; hairs rough, generally ascending to spreading. Inflorescence: in 1s to 3s, 1--5-flowered, dense in fruit; bracts present; pedicel +- 0.5 mm, erect. Flower: calyx circumscissile below middle, cup-like in fruit, hairs +- like leaf hairs; corolla limb generally 1--2 mm diam, appendages inconspicuous, white to pale yellow. Fruit: nutlets +- flattened, base truncate; abaxial ridge 0; adaxial attachment scar edges not raised, abutted near tip.

Greeneocharis circumscissa (Hook. & Arn.) Rydb. var. rosulata (J.T. Howell) Hasenstab & M.G. Simpson
Habit: Plant 0.5--3.5 cm, generally prostrate; root, plant base not red-purple. Stem: 0.5--2 cm, branches few, generally compact; strigose, hairs generally ascending. Leaf: oblanceolate to narrow-oblong, margin +- flat. Flower: calyx 2.5--3 mm in fruit; corolla appendages minute, white. Fruit: nutlets (2)3 or 4, 1.5--1.8 mm, similar, lance-triangular, smooth, shiny, gray, margin a +- flat linear rim, tip acuminate; abaxially +- flat; adaxially biconvex, attachment scar edges occasionally gapped at base; axis not to nutlet tips.
Ecology: Barren granitic gravels; Elevation: 2950--3650 m. Bioregional Distribution: SNH (Inyo, Tulare cos.). Flowering Time: Jul--Aug
Synonyms: Cryptantha circumscissa (Hook. & Arn.) I.M. Johnst. var. rosulata J.T. Howell
Jepson eFlora Author: Ronald B. Kelley, Kristen Hasenstab-Lehman, & Michael G. Simpson
Reference: Johnston 1925 Contr Gray Herbarium 74:1--125; Mathew & Raven 1962 Madroño 16:168--171; Simpson & Hasenstab 2009 Crossosoma 35:1--59; Hasenstab-Lehman & Simpson 2012 Syst Bot 37: 738--757; Simpson et al. 2017 Taxon 66:1406--1420
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)
Listed on CNPS Rare Plant Inventory

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Citation for this treatment: Ronald B. Kelley, Kristen Hasenstab-Lehman, & Michael G. Simpson 2021, Greeneocharis circumscissa var. rosulata, in Jepson Flora Project (eds.) Jepson eFlora, Revision 9,, accessed on June 16, 2024.

Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora,, accessed on June 16, 2024.

No expert verified images found for Greeneocharis circumscissa var. rosulata.

Geographic subdivisions for Greeneocharis circumscissa var. rosulata:
SNH (Inyo, Tulare cos.).
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).


Data provided by the participants of the  Consortium of California Herbaria.
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All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.

CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).