Common Name: WATERLEAF FAMILY Habit: Annual, perennial herb, generally hairy, generally taprooted. Stem: prostrate to erect. Leaf: simple to pinnately compound, basal or cauline, alternate or opposite; stipules 0. Inflorescence: cyme, generally raceme-like and coiled, or flowers 1. Flower: bisexual, generally radial; calyx lobes generally 5, generally fused at base, generally persistent, enlarging in fruit; corolla rotate to cylindric, generally deciduous, lobes generally 5, appendages in pairs on tube between filaments or 0; stamens generally 5, epipetalous, filament base sometimes appendaged, appendages scale-like; ovary generally superior, chambers 1 or 2, placentas 2, parietal, enlarged into chamber, sometimes meeting so ovary appears 2--5-chambered, style 1, style branches 2, stigmas generally head-like. Fruit: capsule, generally loculicidal; valves generally 2. Genera In Family: 12 genera, 240--260 species: especially western US; some cultivated (Emmenanthe, Nemophila, Phacelia). Note: Included in Boraginaceae in TJM2 and some other treatments (Angiosperm Phylogeny Group IV 2016 Bot J Linn Soc 181:1--20) but treated as separate family (excluding Namaceae) by Boraginales Working Group (Luebert et al. 2016). eFlora Treatment Author: Genevieve K. Walden, Robert W. Patterson & Richard R. Halse, except as specified Scientific Editor: Bruce G. Baldwin
Etymology: Howell flower, for J.T. Howell, California botanist, 1903--1994 eFlora Treatment Author: Genevieve K. Walden, Robert W. Patterson, Laura M. Garrison & Debra R. Hansen Reference: Walden & Patterson 2010 Madroño 57:268--273.
Howellanthus dalesianus (J.T. Howell) Walden & R. Patt.
NATIVE Habit: Perennial herb 5--15(20) cm, from +- thick caudex. Leaf: in basal rosette, blade 10--50 mm, +- = petiole, oblong to elliptic, entire. Inflorescence: peduncles axillary, from base, few, decumbent, densely glandular-hairy; leaf-like bracts 0--3, subtending inflorescence; flowers 1--several; pedicel 10--20 mm in fruit. Flower: calyx lobes 3--5 mm, 0.5--1 mm wide in flower, 4--7 mm, 1--1.5 mm wide in fruit, +- not alike, especially in fruit, lanceolate to oblanceolate; corolla 5--10 mm, rotate, white, throat purple-marked, limb 5--15 mm diam, corolla scales free from filaments, semi-circular; stamens (4)6--8 mm, included to slightly exserted, equal, attached at same level, glabrous, filament not widened at base; style (3)6--7 mm, +- included, cleft nearly to base, generally glabrous, occasionally sparsely hairy proximally. Fruit: +- 4 mm, +- spheric, hairy. Seed: 2--4, 2.5--4 mm; surface honeycombed or pitted, attached fleshy structure 0. Chromosomes: n=8. Ecology: Ultramafic substrates, meadows, streambanks, conifer forest; Elevation: 1500--2000 m. Bioregional Distribution: KR. Flowering Time: May--Aug Synonyms: Phacelia dalesiana J.T. Howell Jepson eFlora Author: Genevieve K. Walden, Robert W. Patterson, Laura M. Garrison & Debra R. Hansen Reference: Walden & Patterson 2010 Madroño 57:268--273. Index of California Plant Names (ICPN; linked via the Jepson Online Interchange) Previous taxon: Howellanthus Next taxon: Hydrophyllum
Botanical illustration including Howellanthus dalesianus
Citation for this treatment: Genevieve K. Walden, Robert W. Patterson, Laura M. Garrison & Debra R. Hansen 2021, Howellanthus dalesianus, in Jepson Flora Project (eds.) Jepson eFlora, Revision 9, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=95351, accessed on February 07, 2025.
Citation for the whole project: Jepson Flora Project (eds.) 2025, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on February 07, 2025.
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