Common Name: BORAGE FAMILY Habit: Annual, perennial herb, or shrub, often bristly or sharp-hairy. Stem: prostrate-decumbent to erect. Leaf: basal and/or cauline, simple, generally alternate, sometimes opposite, especially at base. Inflorescence: cymes, arranged singly or in groups of 2--5, generally coiled in flower, generally elongating in fruit. Flower: bisexual, generally radial; sepals 5, free or fused at least at base; corolla 5-lobed, salverform, funnel-shaped, rotate, or bell-shaped, appendages (often called "fornices") 0 or 5 at top of tube, when present often differentially pigmented, alternate stamens; stamens epipetalous; ovary superior, 4-lobed, style 1, entire or minutely 2-lobed (2-branched). Fruit: nutlets 1--4, when > 1, all similar (often called "homomorphic") or 1 or 2 dissimilar in size and/or shape from the others (often called "heteromorphic"), free (fused), smooth to roughened, prickly or bristly or not. Genera In Family: +- 90 genera, +- 1600--1700 species: mostly temperate, especially western North America, Mediterranean; some cultivated (Borago, Echium, Myosotis, Symphytum). Toxicity: Many genera may be TOXIC from pyrrolizidine alkaloids or accumulated nitrates. Note: Sometimes still treated in broader sense of TJM2 (e.g., APG IV 2016 Bot J Linn Soc 181:1--20), but recent evidence (Luebert et al. 2016) supports segregation, for our flora, of the families Ehretiaceae, Heliotropiaceae, Hydrophyllaceae, Lennoaceae, and Namaceae. eFlora Treatment Author: Michael G. Simpson, C. Matt Guilliams, Kristen Hasenstab-Lehman & Ronald B. Kelley Scientific Editor: Bruce G. Baldwin, C. Matt Guilliams, Kristen Hasenstab-Lehman, David J. Keil, Ronald B. Kelley, Robert W. Patterson, Thomas J. Rosatti & Michael G. Simpson
Common Name: STICKSEED Habit: Perennial herb (biennial); hairs appressed to spreading; caudex generally branched in age, often +- woody, taprooted. Stem: ascending or erect. Leaf: lowest petioles generally +- = blades, +- winged, others 0. Inflorescence: coiled cymes, generally > 3, generally terminal and axillary, +- bracted; pedicel in fruit elongated, recurved to reflexed. Flower: calyx deep-5-lobed; corolla rotate-salverform, generally white with yellow patch adaxially, lobes appendaged near base. Fruit: nutlets erect, > style, attachment scar lateral-medial, generally with barb-tipped prickles abaxially and on margin. Etymology: (J. Hackel, Czech botanist, 1783--1869) Note: Values for corolla limb diam take into account shrinkage during flower period. Difficult, study needed, especially in northern California, southeastern Asia; sometimes merged with Lappula. Unabridged Note: In North America filling old-world role of flat-flowered forget-me-nots, Myosotis, Lepechiniella, etc, for pollinators. eFlora Treatment Author: Ronald B. Kelley & Robert L. Carr Unabridged Reference: Gentry & Carr 1976 Mem New York Bot Gard 26:121--227
Hackelia cusickii (Piper) Brand
NATIVE Stem: 1--5 dm, slender; hairs at mid-stem generally strongly appressed upward. Leaf: appressed-stiff-hairy; basal 5--18 cm, 0.4--2.8 cm wide, narrow-elliptic; lower cauline 5--10 cm, 0.4--1.1 cm wide, narrow-elliptic to lanceolate. Inflorescence: open, few-flowered, strigose; pedicel 6--7 mm in fruit. Flower: calyx 4--5 mm; corolla tube generally = calyx, throat open, limb(5)7--13 mm diam, +- blue, appendages +- longer than wide. Fruit: nutlets 3--4.5 mm, abaxial prickles 8--15, < marginal. Chromosomes: 2n=48. Ecology: Under junipers; Elevation: 1300--1950 m. Bioregional Distribution: MP; Distribution Outside California: to east-central Oregon. Flowering Time: May--Jul Jepson eFlora Author: Ronald B. Kelley & Robert L. Carr Index of California Plant Names (ICPN; linked via the Jepson Online Interchange) Listed on CNPS Rare Plant Inventory Previous taxon: Hackelia californica Next taxon: Hackelia floribunda
Botanical illustration including Hackelia cusickii
Citation for this treatment: Ronald B. Kelley & Robert L. Carr 2012, Hackelia cusickii, in Jepson Flora Project (eds.) Jepson eFlora, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=27521, accessed on February 08, 2025.
Citation for the whole project: Jepson Flora Project (eds.) 2025, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on February 08, 2025.
No expert verified images found for Hackelia cusickii.
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