Jepson eFlora: Taxon page
Vascular Plants of California
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Calyptridium parryi var. hesseae

Higher Taxonomy
Family: MontiaceaeView DescriptionDichotomous Key
Habit: Annual to perennial herb; generally fleshy. Stem: 1--many, generally glabrous. Leaf: simple, alternate or opposite. Inflorescence: axillary or terminal; cyme, raceme, panicle, umbel, or flower 1. Flower: bisexual, radial; sepals generally 2(9), free; petals (1)2--19, free or +- fused; stamens 1--many, epipetalous or not, anthers pink, rose, or yellow; ovary superior, chamber 1, ovules 1--many, placenta basal or free-central; styles (0)1--8, generally fused at base, branched. Fruit: capsule, circumscissile or 2--3-valved. Seed: 1--many, shiny or +- pebbly or sculptured, black or gray, generally with oil-filled appendage as food for ants.
Genera In Family: +- 22 genera, +- 230 species: generally temperate America, Asia, Australia, Europe, Kerguelen Is, New Zealand, southern Africa, poorly represented in Europe; some cultivated (Lewisia, Calandrinia). Note: All CA genera previously included in Portulacaceae; details of flowers, seeds require 20× magnification.
Unabridged Note: May include Hectorellaceae.
eFlora Treatment Author: John M. Miller, except as noted
Scientific Editor: Thomas J. Rosatti.
Genus: CalyptridiumView DescriptionDichotomous Key

Common Name: PUSSYPAWS
Habit: Annual, perennial herb, +- fleshy, from taproot or fibrous roots, glabrous. Stem: 1--several, generally spreading to ascending. Leaf: basal or basal and cauline, simple, oblanceolate to spoon-shaped; basal rosetted. Inflorescence: raceme, panicle, or umbel, scapose, bracts generally < sepals, leaf-like or not; flowers generally on 1 side of axis, persistent in fruit or not; pedicels appearing jointed at base with a transverse groove or constriction or not. Flower: sepals 2, ovate to reniform, generally scarious or scarious-margined, persistent in fruit; petals 2--4, < sepals, tips adherent, forming cap in fruit (fruit cap), falling as 1 unit; stamens 1--3, anthers pink, rose, or yellow; style included to exserted, stigmas 2. Fruit: 2-valved, generally compressed, narrowly oblong to +- round, generally translucent, deciduous or not. Seed: 1--many, black, dull, with fine to coarse papillae, to shiny, with papillae 0.
Species In Genus: 9 species: western North America. Etymology: (Greek: cap, for petal tips in fruit) Note: Calyptridium parryi var. arizonicum raised to species rank, as Calyptridium arizonicum, by Simpson et al. (2010) based on morphological and geographic distinctions.
eFlora Treatment Author: C. Matt Guilliams & John M. Miller
Reference: Hershkovitz 2006 Gayana Bot 63:13--74; Simpson et al. 2010 Madroño: 57:145--160
Unabridged Reference: Hershkovitz 1990 Phytologia 68:267--270; Hershkovitz 1991a Ann Missouri Bot Gard 78:1009--1021; Hershkovitz 1991b Phytologia 70:209--225; Hershkovitz 1992 Syst Bot 17:220--238; Hershkovitz & Zimmer 2000 Molec Phylogen Evol 15:419--439; Hinton 1975 Brittonia 27:197--208; Howell 1945 Leafl W Bot 4:214--216; Kelley et al. 2003 FNANM: 4:460--464; Thomas 1956 Leafl W Bot 8:9--11
Species: Calyptridium parryiView Description 

Habit: Annual 2--11 cm; taproot slender. Stem: spreading to ascending, leafy. Leaf: basal and cauline, 1--3 cm, basal generally withering in fruit. Inflorescence: axillary, raceme or panicle, open to dense, 1--3.5 cm; bracts ovate to elliptic; pedicel stout, <= 1 mm, generally appearing jointed at base; flowers persistent in fruit to not. Flower: sepals 2--4.5 mm, generally unequal (outer >, wider than inner), round, ovate, or reniform, generally scarious on margin or becoming scarious throughout; petals 4, 1.5--3 mm, generally white; stamens (1)3; style 0.5--1 mm, stigma +- sessile. Fruit: 3--6 mm, ovate to oblong, deciduous or not. Seed: 5--20, 0.5--0.7(0.8) mm, papillae marginal or throughout. Chromosomes: 2n=44.
Note: Unnamed variety in Baja California.
Calyptridium parryi A. Gray var. hesseae J.H. Thomas
Flower: sepals ovate, margin generally narrowly scarious. Fruit: readily deciduous. Seed: 0.5--0.6 mm, papillae marginal.
Ecology: Sandy soils in chaparral, oak woodland, conifer forest; Elevation: 600--1050 m. Bioregional Distribution: sw SnFrB, n SCoR. Flowering Time: Apr--Jul
Synonyms: Cistanthe parryi var. hesseae (J.H. Thomas) Kartesz & Gandhi
Jepson eFlora Author: C. Matt Guilliams & John M. Miller
Reference: Hershkovitz 2006 Gayana Bot 63:13--74; Simpson et al. 2010 Madroño: 57:145--160
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)
Listed on CNPS Rare Plant Inventory

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Botanical illustration including Calyptridium parryi var. hesseae

botanical illustration including Calyptridium parryi var. hesseae


Citation for this treatment: C. Matt Guilliams & John M. Miller 2014, Calyptridium parryi var. hesseae, in Jepson Flora Project (eds.) Jepson eFlora, Revision 2,, accessed on June 26, 2022.

Citation for the whole project: Jepson Flora Project (eds.) 2022, Jepson eFlora,, accessed on June 26, 2022.

Calyptridium parryi var. hesseae
click for enlargement
© 2017 Chris Winchell
Calyptridium parryi var. hesseae
click for enlargement
© 2020 Dylan Neubauer
Calyptridium parryi var. hesseae
click for enlargement
© 2017 Chris Winchell
Calyptridium parryi var. hesseae
click for enlargement
© 2017 Chris Winchell
Calyptridium parryi var. hesseae
click for enlargement
© 2017 Chris Winchell
Calyptridium parryi var. hesseae
click for enlargement
© 2017 Chris Winchell

More photos of Calyptridium parryi var. hesseae in CalPhotos

Geographic subdivisions for Calyptridium parryi var. hesseae:
sw SnFrB, n SCoR.
1. You can change the display of the base map layer control box in the upper right-hand corner.
2. County and Jepson Region polygons can be turned off and on using the check boxes.
map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).


Data provided by the participants of the  Consortium of California Herbaria.
View all CCH records
All markers link to CCH specimen records. The original determination is shown in the popup window.
Blue markers indicate specimens that map to one of the expected Jepson geographic subdivisions (see left map). Purple markers indicate specimens collected from a garden, greenhouse, or other non-wild location.
Yellow markers indicate records that may provide evidence for eFlora range revision or may have georeferencing or identification issues.

CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).