University Herbarium
		University of California, Berkeley

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	photo by Game

Plants erect in glossy red-brown tufts (sometimes rich green in shaded sites), to 6 cm high. Leaves evenly inserted along stem, not at all comose. Leaves somewhat concave and keeled, elliptic to elliptic-lanceolate, broadest near leaf middle, bluntly rounded to cucullate at apex, to 3 mm long, about 3: 1, erect when moist, not much changed when dry, not plicate. Median laminal cells 3–4: 1, elongate-hexagonal with rather thick walls (lumen: wall ratio 4–6: 1), completely smooth, not pitted, to 18 µm wide. Basal and alar cells to 25 µm broad, quadrate to short rectangular, 1–1.5: 1, in an abruptly defined patch, usually with bistratose areas. Margins not decurrent, erect throughout and nearly entire. Limbidium essentially absent but marginal cells somewhat elongate and narrowed. Costa reddened, especially at base, ending only a few cells below apex or percurrent. Costa cross-section elliptic, about 2: 1, with a well-demarcated abaxial stereid band and with two layers of rather small cells adaxial to the stereid band. Cells of axillary hairs almost uniform throughout, to 6 cells long, to 200 µm, with basal cells unpigmented and almost as long as the distal cells. Micronemata absent. Macronemata mostly smooth or with smaller branches papillose, to 30 µm wide at insertion, pale brown, closely branched, surrounding an isodiametric macronematal area. Stem cross-section rounded-pentagonal but ribbed by costa bases, with a strongly demarcated central strand, with rather pachydermous corticals and with an outer stereome only 2–3 cells thick between the ridges formed by the costa bases, never with an hyaloderm. Gemmae absent from our specimens.
    Dioicous with male plants somewhat smaller than females. Perichaetial leaves similar but smaller relative to the vegetative leaves. Seta red-brown to brown, smooth, erect but geniculate distally, to 20 mm long. Urn with neck comprising about 1/4–1/3 of capsule length, to 3.5 mm long, cylindric, 3.5–4.5: 1, pendulose to pendent, neither sulcate nor strumose. Operculum short conic. Annulus inconspicuous, falling in fragments. Suboral exothecial cells to 25 µm broad, transversely elongate in several rows. Exothecial cells at middle of urn irregularly rectangular with curved lateral walls, 2–4: 1, to 20 µm broad, not in regular rows, evenly thick walled with lumen: wall ratio 2–4: 1. Stomates restricted to neck of capsule, not raised on pustules, elongate. Exostome yellow-brown, to 600 µm long, obviously shouldered, lightly and evenly punctate-papillose. Endostome with basal membrane constituting about 1/2 of total length of endostome. Segments smooth to lightly papillose, somewhat shorter than exostome, perforated as orbicular holes along keel, with 2–4 appendiculate cilia. Calyptra cucullate, mostly shed before capsule maturity. Spores to 18 µm, finely papillose.

Elevation by latitude plot for Bryum miniatum
   in California

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