The Jepson Manual: Higher Plants of California

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Names are linked to treatments from the Manual


Key to CARDAMINE

N.B.: couplets are linked, e.g., 1. is linked to 1'.

  1. Leaves simple, shallowly lobed to generally entire, rarely ± dissected
 
    2. Plant ± scapose, rarely with 1–2 cauline leaves, 2–12 cm; basal leaves many, blade base ± tapered; inflorescence in fruit ± umbel-like
C. bellidifolia var. pachyphylla
    2' Plant with cauline leaves, 15–50 cm; basal leaves few, blade base abruptly narrowed; inflorescence in fruit elongate
 
      3. Cauline leaves from base to inflorescence; rhizome leaves 0; stems amid generally dense rhizomes
C. cordifolia var. lyallii
      3' Cauline leaves near inflorescence; rhizome leaves present; stems from ends of elongate rhizomes
C. pachystigma
        4. Cauline leaves 1-palmately or -pinnately ± dissected
var. dissectifolia
        4' Cauline leaves entire or shallowly lobed
var. pachystigma
  1' Leaves or at least lower cauline palmately or pinnately lobed to compound
 
    5. Stem widened to strong attachment with rhizome or rhizome 0; rhizome leaves generally 0; cauline leaves pinnately lobed to compound
 
      6. Annual or biennial from fibrous roots or weak taproot
 
        7. Basal leaves 0–few, not rosetted, generally deciduous; lower stem hairs few or generally 0
C. pensylvanica
        7' Basal leaves many, generally ± rosetted, persistent; lower stem hairs 0–many
 
          8. Stems 1–many, erect, 5–30 cm, outer 0 or decumbent; fruit < 1 mm wide; alien
C. hirsuta
          8' Stems 1–several, erect to ascending, generally > 20 cm; fruit 1–2 mm wide; native
C. oligosperma
      6' Perennial from rhizomes or strong taproot
 
        9. Petals 8–14 mm; style 1.5–6 mm
 
          10. Rhizome elongate, > 3 cm, not tuber-like, ± spreading; style 1.5–3(4) mm; leaflets widely dentate to lobed
C. angulata
          10' Rhizome short, < 2 cm, tuber-like, ± spreading to ± erect; style 3–6 mm; leaflets or lobes entire to wavy or dentate
2 C. californica (see 17. for vars.)
        9' Petals 4–6 mm; style ± 1–1.2 mm
 
          11. Basal leaves pinnately lobed; rhizome ± tuber-like at stem base
C. occidentalis
          11' Basal leaves generally entire to shallowly palmately lobed; rhizome not tuber-like at stem base
C. breweri
            12. Leaflets of lower cauline leaves wedge-shaped to truncate at base; rhizome ± elongate, spreading
var. breweri
            12' Leaflets of lower cauline leaves cordate at base; rhizome generally short, ± erect
var. orbicularis
    5' Stem narrowed to weak attachment with rhizome; rhizome leaves present; cauline leaves pinnately or generally palmately lobed to compound
 
      13. Stem slender, < 2 dm; rhizome ovoid to elongate, generally with ovoid, tuber-like swelling from which leaf arises, 0.5–3 cm; inflorescence in fruit ± umbel-like
C. nuttallii
        14. Rhizome leaf simple, ± entire to shallowly lobed
var. nuttallii
        14' Rhizome leaf subpalmately compound, leaflets 3–5, ± entire to ± lobed
 
          15. Rhizome leaflets (3)5, oblong, ± entire, 6–10 mm wide
var. covilleana
          15' Rhizome leaflets 3–5, ovate to ± elliptic to obovate, toothed or lobed, 10–20 mm wide
 
            16. Rhizome leaflets ± sessile, toothed or lobed; rhizome elongate, generally whitish
var. dissecta
            16' Rhizome leaflets stalked, toothed; rhizome ovoid to slightly elongate, generally yellowish
var. gemmata
      13' Stem ± stout, 2–7 dm; rhizome ovoid to subspheric, tuber-like, < 2 cm; inflorescence in fruit elongate
2 C. californica
        17. Rhizome leaves 1–2-pinnate, leaflets 5–9, often lobed
var. cuneata
        17' Rhizome leaves simple, entire, or leaflets 3, rarely 5 and ± pinnately arranged
 
          18. Rhizome leaves generally with 3 leaflets, rarely with 5 or simple, entire
 
            19. Leaflets or lobes of cauline leaves generally ovate or wider, thin, generally sharp-toothed
var. californica
            19' Leaflets or lobes of cauline leaves generally oblong to linear, thickish, generally entire or teeth small, few or only 1 at tip
var. integrifolia
          18' Rhizome leaves simple
 
            20. Cauline leaves simple; rhizome leaf blade margin shallowly wavy
var. cardiophylla
            20' Cauline leaves generally compound; rhizome leaf blade margin wavy to deeply dentate
var. sinuata


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