| 1. Involucral bracts and awns (or ribs) 35 |
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| 2. Involucral bracts and awns 3 |
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| 3. Involucral tube cylindric, prominently cross-ridged, glabrous; stamens 6; Desert |
| C. corrugata |
| 3' Involucral tube 3-angled, ± bell-shaped or flaring, smooth to moderately cross-ridged, hairy; stamens 9; Southwestern California, East of Sierra Nevada, Desert |
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| 4. Plant erect; awns of inflorescence and involucral bracts straight East of Sierra Nevada, Desert |
| C. rigida |
| 4' Plant prostrate to decumbent; inflorescence bracts unawned, involucral awns hooked |
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| 5. Involucral tube very narrowly flaring, smooth, ± brown; South Coast |
| C. orcuttiana |
| 5' Involucral tube bell-shaped, thinly hairy, reddish; Peninsular Ranges |
| 2 C. polygonoides var. longispina |
| 2' Involucral bracts and awns 45 |
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| 6. Awns straight; perianth 2.53.5 mm, glabrous; stamens attached at base of perianth tube; Mojave Desert |
| C. spinosa |
| 6' Awns hooked; perianth 1.52.5 mm, hairy; stamens attached near top of perianth tube |
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| 7. Involucral tube cylindric, 34.5 mm, largest bract ± leaf-like; perianth yellow, thinly hairy; Great Basin Floristic Province, Mojave Desert, n Western Transverse Ranges |
| C. watsonii |
| 7' Involucral tube bell-shaped, 1.52.5 mm, bracts not leaf-like; perianth white to rose, densely hairy; California Floristic Province |
| C. polygonoides |
| 8. Involucre (including awn) 34 mm; Peninsular Ranges |
| var. longispina |
| 8' Involucre (including awn) ± 5 mm; n California |
| var. polygonoides |
| 1' Involucral bracts and awns (or ribs) 6 |
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| 9. Involucral bract margin scarious or membranous |
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| 10. Involucral bract margin membranous, continuous across sinuses (involucre thus mimicking a fused corolla); true cauline leaves (no inflorescence or involucre in axil) generally present; generally inland California Floristic Province |
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| 11. Cauline leaves (and lower bracts) alternate; involucre ± woolly or becoming glabrous, bracts ± equal; perianth 1.53 mm, densely hairy California Floristic Province |
| C. membranacea |
| 11' Whorl of true cauline leaves present near mid-stem in most plants; involucre bristly, bracts equal or not; perianth 3.55 mm, sparsely hairy |
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| 12. 3 involucral bracts >> alternating 3, membrane purple; South Coast Ranges |
| C. douglasii |
| 12' 6 involucral bracts ± equal, membrane white; foothills around n&e Great Central Valley |
| 2 C. stellulata |
| 10' Involucral bract margin scarious, not continuous across sinuses (except sometimes between pairs of erect bracts); true cauline leaves generally 0; ± coastal Central Western California, North Coast (except C. stellulata) (C. pungens complex) |
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| 13. Involucral awns straight |
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| 14. Plant generally decumbent; involucral bracts spreading, awns brown; perianth 34.5 mm; c North Coast |
| C. howellii |
| 14' Plant generally ± erect; involucral bracts erect, awns ivory or straw-colored; perianth 46 mm; n Central Coast |
| 2 C. valida |
| 13' Involucral awns hooked |
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| 15. Perianth glabrous, lobes white, tube yellow |
| C. diffusa |
| 15' Perianth hairy, lobes same color as tube |
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| 16. Plant generally erect; whorl of true cauline leaves often present; involucral tube 34 mm, bracts clearly margined |
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| 17. Plant decumbent to erect, soft-hairy; involucre thinly hairy; perianth 2.54 mm; n Central Coast, s San Francisco Bay Area |
| C. robusta |
| 17' Plant strictly erect, coarse-hairy; involucre bristly; perianth 45 mm; foothills around n Great Central Valley |
| 2 C. stellulata |
| 16' Plant prostrate to ascending; cauline leaves 0; involucral tube 1.53 mm, bract margins sometimes indistinct |
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| 18. Involucral bracts obviously white- to purple-margined, tube 23 mm; stamens 9 |
| C. pungens |
| 18' Involucral bract margins thin, ± indistinct; involucral tube 1.52.5 mm; stamens 39 |
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| 19. Perianth lobes barely exserted, entire to jagged (central tooth, if any, generally not greatly predominant); stamens 39; s Central Coast |
| 2 C. angustifolia |
| 19' Perianth lobes well exserted, with an abrupt central tooth (if teeth > 1, central greatly predominant); stamens 9; n Central Coast, San Francisco Bay Area |
| 2 C. cuspidata |
| 9' Involucral bract margin not scarious or membranous |
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| 20. Involucral awns unequal, longest generally > 2 × length of next longest in some or all involucres |
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| 21. Longest involucral awn hooked, ± 2 × length of next longest Northwestern California, Central Western California, s Sierra Nevada |
| C. clevelandii |
| 21' Longest involucral awn straight, generally 35 × length of next longest |
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| 22. Longest involucral awn ± 5 × length of next longest eastern Central Western California, s Sierra Nevada, n Western Transverse Ranges |
| C. uniaristata |
| 22' Longest involucral awn ± 3 × length of next longest |
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| 23. Involucres (including awns) generally 57 mm, scattered in small clusters; Outer South Coast Ranges |
| C. rectispina |
| 23' Involucres generally 810 mm, clumped in large clusters; Inner South Coast Ranges |
| C. ventricosa |
| 20' Involucral awns equal or of alternating lengths, longest generally < 2 × length of next longest |
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| 24. Inner perianth lobes either rather deeply fringed or notched (may require dissection of involucre to see) |
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| 25. Outer perianth lobes fringed along sides; Southwestern California |
| C. fimbriata |
| 26. Fringe segments, especially lobe tip, wide, ± lobe-like; s South Coast, w Peninsular Ranges |
| var. fimbriata |
| 26' Fringe segments, including lobe tip, thin, not lobe-like; e Peninsular Ranges |
| var. laciniata |
| 25' Outer perianth lobes entire or tip notched or fringed; South Coast Ranges (C. palmeri complex) |
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| 27. Perianth lobes white to pinkish |
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| 28. Inner perianth lobes notched, 56 mm; plant yellowish green; Western Transverse Ranges (Sierra Madre Mtns, Santa Barbara Co.) |
| C. blakleyi |
| 28' Inner perianth lobes fringed, 45 mm; plant reddish or greenish; Outer South Coast Ranges |
| C. obovata |
| 27' Perianth lobes red to purple |
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| 29. Outer perianth lobes entire, erect; on serpentine; Outer South Coast Ranges |
| C. palmeri |
| 29' Outer perianth lobes ± notched, spreading to recurved; not on serpentine; South Coast Ranges |
| C. biloba |
| 30. Outer perianth lobes deeply notched; South Coast Ranges |
| var. biloba |
| 30' Outer perianth lobes shallowly notched; Inner South Coast Ranges |
| var. immemora |
| 24' Inner perianth lobes entire, with a central tooth, or shallowly jagged (may require dissection to see) |
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| 31. Perianth lobes all equal in length and width |
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| 32. Plant erect; inflorescence breaking at nodes; stamens 3, attached near top of perianth tube |
| C. brevicornu |
| 33. Leaves ± linear; generally Desert |
| var. brevicornu |
| 33' Leaves oblanceolate; East of Sierra Nevada |
| var. spathulata |
| 32' Plant prostrate to decumbent; inflorescence not breaking at nodes; stamens 39, attached at base of perianth tube; w California Floristic Province |
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| 34. Perianth lobes entire; plant thinly hairy, yellowish green when fresh; filaments fused at base; Southwestern California |
| C. procumbens |
| 34' Perianth lobes ± jagged or with a central tooth; plant densely hairy, brown or gray; filaments free; Central Coast |
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| 35. Perianth lobes barely exserted, entire to jagged (central tooth, if any, generally not greatly predominant); stamens 69; s Central Coast |
| 2 C. angustifolia |
| 35' Perianth lobes well exserted, with an abrupt central tooth (if teeth > 1, central greatly predominant); stamens 9; n Central Coast, San Francisco Bay Area |
| 2 C. cuspidata |
| 31' Perianth lobes unequal, inner generally < and narrow than outer |
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| 36. Most involucral awns straight; perianth lobes sometimes ± jagged at tip |
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| 37. Plant decumbent; involucre 1.52 mm, bracts spreading; South Coast |
| C. parryi var. fernandina |
| 37' Plant ± erect; involucral tube 34.5 mm, bracts erect; s North Coast |
| 2 C. valida |
| 36' Involucral awns hooked; perianth lobes generally entire |
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| 38. Plant grayish, canescent; involucral tube 1.52 mm Transverse Ranges |
| C. parryi var. parryi |
| 38' Plant reddish or greenish, thinly hairy; involucral tube 2.56 mm (C. staticoides complex) |
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| 39. Lower bracts ± linear, not very leaf-like, deciduous |
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| 40. Perianth well exserted; involucres loosely clustered; eastern Peninsular Ranges |
| C. leptotheca |
| 40' Perianth moderately exserted; involucres densely clustered; s Central Western California, Southwestern California |
| C. staticoides |
| 39' Lower bracts ± oblanceolate, ± leaf-like, persistent |
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| 41. Involucral tube 3.55 mm, cylindric; not on serpentine |
| C. xanti |
| 42. Involucre densely hairy; eastern San Bernardino Mountains, n San Jacinto Mountains |
| var. leucotheca |
| 42' Involucre thinly hairy; s Sierra Nevada Foothills, Inner South Coast Ranges, n border Transverse Ranges, East of Sierra Nevada |
| var. xanti |
| 41' Involucral tube generally 23 mm, ± bell-shaped; on serpentine or not |
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| 43. Inflorescence clusters ± open; involucral tube generally 2.53 mm; stamens 9; serpentine; s Outer South Coast Ranges |
| C. breweri |
| 43' Inflorescence clusters head-like; involucral tube generally 2-2.5 mm; stamens 6; non-serpentine; n Channel Islands |
| C. wheeleri
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