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Alan R. Smith and Thomas Lemieux

Perennial, in soil or on or among rocks; rhizome creeping to erect, scaly
Leaves generally all ± alike (or of 2 kinds, fertile and sterile), generally < 50 cm, often < 25 cm; petiole generally thin, wiry, often dark, in transverse section with vascular strands generally 1–3, less often many in a circle; blade generally pinnate or ± palmate-pinnate (see Adiantum ), often 2 or more compound, lower surface often with glands, ± powdery exudate, hairs, or scales; segments round, oblong, fan-shaped, or otherwise, veins generally free
Sporangia in sori or not, marginal, submarginal, or along veins, sometimes covered by recurved, often modified segment margins (false indusia); true indusia 0; spores spheric, sides sometimes flat, scar with 3 radiating branches
Genera in family: ± 40 genera, 500 species: worldwide, especially dry areas. Definition of Cheilanthes and related genera problematic; traditional limits often untenable.


Plant in soil or rock crevices; rhizome short- to long-creeping-decumbent, generally many-branched, scales generally linear-lanceolate, pale to dark, with darker mid-stripe or not
Leaf < 75 cm; petiole cylindric, reddish brown to blackish; blade generally 2–3-pinnate, generally oblong to narrowly triangular; segments generally small, ± flat or lower side concave (from recurved margins)
Sporangia along margin, in discrete patches to continuous, partly to completely covered by recurved margin (generally not recurved in C. cooperae)
Species in genus: 150+ species: generally Am, generally dry areas
Etymology: (Greek: lip flower, from location of sporangia)
Horticultural information: DFCLT.


C. parryi (D.C. Eaton) Domin

Rhizome short-creeping, > 6 cm; scales medium brown, most with darker (thread-like) mid-stripe
Leaf 6–15(25) cm, 1–2(3) cm wide; scales 0; petiole < 1 mm wide, hairs short to long, bent, appressed to ± spreading, glandular and not, pale; segments small, ± round, ± flat, hairs long (4+ mm), tangled, generally non-glandular, very dense on both surfaces, making upper surface silver-whitish, lower tan to brown or golden
Sporangia ± visible through hairs at segment margin; spores blackish
Chromosomes: 2n=60
Ecology: Limestone, granite crevices, rocks
Elevation: 100–1500 m.
Bioregional distribution: Peninsular Ranges, East of Sierra Nevada, Desert
Distribution outside California: to Utah, Arizona, Baja California

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