TREATMENT FROM THE JEPSON MANUAL (1993) previous taxon | next taxon
Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California
For up-to-date information about California vascular plants, visit the Jepson eFlora.

  • Up-to-date information about California vascular plants is available from the Jepson eFlora.



Sherry Whitmore and Alan R. Smith

Perennial, on plants, rocks, in rock crevices, less often in soil, humus, or on dunes; rhizome short- to long-creeping, branched, glaucous to not, scaly
Leaves ± alike or of 2 kinds, fertile and sterile; petiole thin to thick, straw-colored or green to brown or black, generally jointed to persistent knob on rhizome; blade generally simple to 1-pinnate, membranous to fleshy or leathery; veins free to fused, generally forked
Sporangia: sori round to elongate, rarely linear, generally 1 per areole, in 1–several rows on each side of segment midrib; indusium 0; spores generally ± elliptic, ± smooth to coarse-tubercled or -ridged, scar linear
Genera in family: ± 46 genera, ± 650 species: worldwide, especially Old World tropical. Numbers of genera, species depend on treatment; many species cultivated.



Sherry Whitmore

Rhizome long-creeping; scales lanceolate, ± brownish, 1-colored or often with darker central area or midstripe
Leaves 0.2–10(20) dm, ± alike or fertile > sterile; petiole glabrous to scaly; blade 1-pinnate to generally deeply pinnately lobed, rarely simple and unlobed, glabrous to hairy, glandular or not, scales on lower surface midrib near base generally lanceolate or linear-lanceolate, generally ± brown; veins free to fused
Sporangia: sori in 1 row on each side of segment midrib, generally raised, sometimes including branched or unbranched, glandular hairs, sporangium-like structures, or shriveled sporangia; spores yellow
Species in genus: ± 160 species: generally New World, tropical, some temp, few boreal
Etymology: (Latin: many feet, from rhizome)
Reference: [Whitmore & Smith 1991 Madroño 38:233–248]
50% or more malformed spores indicates hybrid involving 2 or more species in CA.


P. calirhiza S.A. Whitmore & A.R. Sm.

Rhizome 5–10 mm diam, ± glaucous or not, taste acrid and ± sweet; scales 1-colored
Leaves ± summer-deciduous if conditions dry; blade (5)10–20(40) cm, oblong-ovate, ± membranous to ± firm, upper sides of midribs hairy, segments serrate, tips generally acute to obtuse, veins free to 35(–50)% fused
Sporangia: sori 1.5–4 mm, generally larger, sometimes fused near segment base, ovate, sometimes with short, many-branched, glandular hairs
Chromosomes: 2n=148
Ecology: On plants, rocky cliffs or outcrops, roadcuts, often granitic or volcanic, rarely dunes
Elevation: < 1220 m.
Bioregional distribution: Northwestern California (except High North Coast Ranges), Cascade Range Foothills, Sierra Nevada (except s Sierra Nevada Foothills), Great Central Valley, Central Western California
Distribution outside California: to Oregon, s Mexico
Hybrids with P. glycyrrhiza sterile, 2n=111, abundant, of similar geography.

previous taxon | next taxon
bioregional map for POLYPODIUM%20calirhiza being generated

Retrieve Jepson Interchange Index to Plant Names entry for Polypodium calirhiza
Retrieve dichotomous key for Polypodium
Return to treatment index page
University & Jepson Herbaria Home Page | Copyright © by the Regents of the University of California