ROSACEAE

ROSE FAMILY

Annual to tree
Leaves simple to pinnately to palmately compound, generally alternate; stipules free to fused, persistent to deciduous
Inflorescence: cyme, raceme, panicle, or flowers solitary
Flower generally bisexual, radial; hypanthium free or fused to ovary, saucer- to funnel-shaped, often with bractlets alternate with sepals; sepals generally 5; petals generally 5, free; stamens (0)5–many, pistils (0)1–many, simple or compound; ovary superior to inferior, styles 1–5
Fruit: achene, follicle, drupe, pome, or blackberry- to raspberry-like
Seeds generally 1–5
Genera in family: 110 genera, ± 3000 species: worldwide, especially temp. Many cultivated for ornamental and fruit, especially Cotoneaster , Fragaria , Malus , Prunus , Pyracantha, Rosa , and Rubus
Reference: [Robertson 1974 J Arnold Arbor 55:303–332,344–401,611–662]
Family description, key to genera by Barbara Ertter and Dieter H. Wilken.

PHYSOCARPUS

NINEBARK

Thomas J. Rosatti

Shrub, generally ± stellate-hairy
Leaves stipuled, petioled, deciduous; blade ovate to ± round in CA, generally palmately 3–7-lobed, lobes crenate to serrate
Inflorescence ± umbel-like, bracted
Flower: hypanthium bell-shaped, bractlets 0; sepals persistent; petals rounded, white; stamens 20–40; pistils 1–5, free or ± fused, ovary superior, style thread-like, stigma head-like
Fruit: follicles, inflated, often opening along both sutures
Seeds 2–4, ovoid; coat hard, shiny
Species in genus: ± 10 species: North America, Asia
Etymology: (Greek: bladdery fruit)

Native

P. capitatus (Pursh) Kuntze

Plant 10–25 dm
Leaf: petiole 5–20+ mm; blade 30–140 mm, often ± glabrous; lobes generally 3–5, generally serrate (if 0, leaf coarsely ± 9–11-toothed)
Flower: sepals 2.5–3 mm; petals 2.5–3 mm
Fruit glabrous to ± hairy
Chromosomes: n=9
Ecology: Moist banks, n-facing slopes, coniferous forests
Elevation: < 1400 m.
Bioregional distribution: California Floristic Province (except Great Central Valley)
Distribution outside California: to Alaska, Montana, Utah
Horticultural information: 4, 5, 6, 17 &IRR, partSHD: 1, 2, 7, 8, 9, 14, 15, 16, 18, 19, 20, 21, 22, 23, 24; CVS: deciduous GRCVR.