TREATMENT FROM THE JEPSON MANUAL (1993) previous taxon | next taxon
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©Copyright 1993 by the Regents of the University of California
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David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.


Annual, perennial herb, subshrubs
Stems erect
Leaves simple or pinnate, opposite or alternate, dotted with embedded oil glands
Inflorescence: heads radiate or discoid, solitary or in few-headed, often leafy-bracted, often open cymes; peduncles slender, with leaf-like or scale-like bracts; involucre bell-shaped to cylindric; phyllaries in 3 series, outer << others, free, 2 inner ± equal, fused or free, gland-dotted; receptacle flat, bearing short, fringed scales
Ray flowers 0–few; corollas yellow to red
Disk flowers few–many; corollas yellow to orange; style tips tapered
Fruit obconic, ribbed; pappus of scales dissected into slender bristles
Species in genus: 10 species: sw US, n Mex
Etymology: (Greek: gland-leaf)
Reference: [Strother 1986 Sida 11:371–378]


A. porophylloides (A. Gray) Strother

Subshrub, glabrous; odor unpleasant
Stems many, 2–6 dm, generally much-branched
Leaves opposite below, alternate above, 1.5–4 cm, deeply divided; lobes linear, entire to sharply serrate, each with 1 gland at base, 1 near tip
Inflorescence: heads radiate (discoid); peduncles 2–8 cm, naked or with 1–5 narrow bracts; involucre cylindric; outer phyllaries 12–16, 3–8 mm, erect or recurved, each with a central gland; inner phyllaries 12–20, 10–15 mm, lanceolate, fused below, gland-dotted
Ray flowers (0)8–12; ligules 2–4 mm, yellow to red-orange
Disk flowers many; corollas 7–8 mm, yellow-orange
Fruit 5 mm, sparsely hairy; pappus scales 8–12, 7–8 mm, each dissected into 7–11 bristles
Chromosomes: 2n=26
Ecology: Dry, rocky hillsides, washes
Elevation: 200–1460 m.
Bioregional distribution: s Mojave Desert, Sonoran Desert
Distribution outside California: to Arizona, n Mexico
Flowering time: Mar–Jun
Synonyms: Dyssodia p. A. Gray

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