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©Copyright 1993 by the Regents of the University of California
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David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.



John S. Mooring and Dale E. Johnson

Annual to shrubs, ± woolly
Leaves generally alternate, entire to nearly compound
Inflorescence: heads 1–many, generally radiate; cluster often ± flat-topped; involucre obconic to hemispheric; phyllaries in 1 series, free or ± fused; receptacle flat to columnar, generally naked
Ray flowers generally ± 1 per phyllary; ligules entire to lobed, generally yellow (white)
Disk flowers (3)10–300; corolla yellow; anther tips ovate, deltate or awl-shaped
Fruit 4-angled or flattened in outer flowers, generally club-shaped in inner flowers; pappus 0–15 ± jagged or fringed scales
Species in genus: 14 species: w North America
Etymology: (Greek: woolly leaf)
Reference: [Mooring 1991 Madroño 38:213–226]
Annual species by Dale E. Johnson.


E. lanosum (A. Gray) A. Gray

Annual 1–15 cm, decumbent-ascending, often reddish, sparsely woolly
Leaf 5–20 mm, linear-oblanceolate, entire or lobed at tip
Inflorescence: head 1; peduncles 1–5 cm; involucre 5–7 mm, ± cylindric; phyllaries 8–10, acuminate, free; receptacle conic
Ray flowers 8–10; ligules 3–7 mm, oblong, white, sometimes red-veined
Disk flowers many; corollas 2–3 mm, glabrous; anther tips awl-like, glabrous
Fruit 2.5–4.5 mm, linear or narrowly club-like, glabrous to minutely strigose; pappus scales 0.5–2.5 mm, generally very unequal, longest awned
Chromosomes: 2n=8
Ecology: Desert scrub
Elevation: < 1400 m.
Bioregional distribution: Desert
Distribution outside California: to sw Utah, Arizona, nw Mexico
Flowering time: Feb–May
Horticultural information: TRY.

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