BORAGINACEAE

BORAGE FAMILY

Annual, perennial herb, shrubs, generally bristly or sharply hairy
Stem prostrate to erect
Leaves cauline, often with basal rosette, generally simple, alternate; lower sometimes opposite, entire
Inflorescence: cyme, generally elongate, panicle-, raceme- or spike-like, coiled in flower, generally uncoiled in fruit or flowers 1–2 per axil
Flowers generally bisexual, generally radial; sepals 5, free or fused in lower half; corolla 5-lobed, generally salverform, top of tube generally appendaged, appendages 5, alternating with stamens, sometimes arching over tube; stamens 5, epipetalous; ovary superior, generally 4-lobed, style generally entire
Fruit: nutlets 1–4, smooth to variously roughened, sometimes prickly or bristled
Genera in family: ± 100 genera, ± 2000 species: tropical, temp, especially w North America, Medit; some cultivated (Borago, Echium, Myosotis, Symphytum ). Almost all genera may be TOXIC from alkaloids or accumulated nitrates
Recent taxonomic note: Recently treated to include Hydrophyllaceae [Olmsted et al. 2000 Mol Phylog Evol 16:96–112]
Family description, key to genera by Timothy C. Messick.

MYOSOTIS

FORGET-ME-NOT

Elaine Joyal

Annual or perennial herb, glabrous to rough-hairy; roots generally diffuse
Stem decumbent to erect
Leaves: basal generally oblong or oblanceolate; cauline generally linear to elliptic
Inflorescence coiled, generally raceme-like, eventually ± open; bracts 0 (or leaf-like in lower half); flowers generally in upper half of plant
Flower: calyx lobes 5, generally ± equal; corolla salverform or widely funnel-shaped, white, yellow, or generally blue, tube 5-appendaged at top, limb abruptly spreading, 5-lobed; stamens 5, included; style generally included
Fruit: nutlets smooth, shiny, each with raised outer margin, scar lateral, at base, small
Species in genus: 50 species: temp, boreal
Etymology: (Greek: mouse ear, from leaf)
Reference: [Grau 1964 Osterr Bot Zeitschr 111:561–617]
Generally flowers in spring.

Introduced

M. latifolia Poir.

Perennial
Stem < 70 cm; base woody
Leaves: basal large, ovate; cauline oblong
Inflorescence bracted at base; pedicel in fruit ascending to spreading, at least 5 mm, > or = calyx
Flower: calyx 3–6 mm in fruit, tube hairs generally hooked; corolla blue or pink, tube 2 X calyx, limb 5–10 mm diam
Fruit 2–3 mm, > style, widely ovate, dark brown
Chromosomes: 2n=18
Ecology: Moist, disturbed, shady places
Elevation: 0–300 m.
Bioregional distribution: Scattered in California Floristic Province
Distribution outside California: native to nw Africa