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DRYOPTERIDACEAE

WOOD FERN FAMILY

Alan R. Smith and Thomas Lemieux

Perennial, in soil or rock crevices; rhizome generally short-creeping, suberect, or erect, scales large, generally tan to brown, generally 1-colored
Leaves generally tufted, 5–200+ cm, generally ± alike; petiole generally firm, base generally darker, with 2–many vascular strands; blade 1–4-pinnate, often with scales, hair-like scales, hairs (except clear, needle-like hairs generally 0), or short-stalked glands on axes, sometimes between veins, veins free to netted; 1° and 2° axes generally grooved on upper side
Sporangia: sori round, less often oblong or J-shaped, along or at tips of veins; indusia peltate, round-reniform, oblong to linear, J-shaped, hood-like, or cup-like, rarely 0; spores elliptic, winged, ridged, or spiny, scar linear
Genera in family: ± 60 genera, > 1000 species: worldwide, especially tropical, wooded areas. Woodsia sometimes in Woodsiaceae; Athyrium, Cystopteris sometimes in Athyriaceae.

POLYSTICHUM

SWORD FERN

Rhizome generally suberect to erect, often stout
Leaf: petiole stout, firm, generally densely scaly, in X -section with many round vascular strands in an arc; blade generally 1–3-pinnate, thin to leathery, scaly, veins generally free, rarely casually joined; 1° leaflet bases often wider on distal side; teeth generally including bristle-like tips that are < 2 mm
Sporangia: sori round; indusium generally peltate, sinus 0
Species in genus: ± 175+ species: ± worldwide
Etymology: (Greek: many rows, from rows of sori on type sp.)

Native

P. lemmonii Underw.


Leaf 15–40 cm; petiole 1/4–1/3 blade length, base scales ± 2 mm wide, generally linear-lanceolate; blade narrow-lanceolate, 2-pinnate; 1° leaflets generally 1–3 cm; teeth without bristle-like tips
Sporangia: indusium entire or minutely toothed
Chromosomes: 2n=82
Ecology: Serpentine, rocks, ledges, rocky streambeds
Elevation: 1200–2600 m.
Bioregional distribution: Klamath Ranges, High North Coast Ranges, High Cascade Range, n High Sierra Nevada
Distribution outside California: to British Columbia
Horticultural information: DRN: 4, 5 &IRR: 1, 2, 3, 6 &SHD: 7, 15, 16, 17; DFCLT.

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