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BRASSICACEAE

MUSTARD FAMILY

Reed C. Rollins, except as specified

Annual to subshrub
Leaves generally basal and cauline, alternate, generally simple; stipules 0
Inflorescence: generally raceme
Flower bisexual; sepals 4, free; petals (0)4, free, generally white or yellow, often clawed; stamens generally (2,4)6, generally 4 long, 2 short; ovary 1, superior, chambers generally 2, septum membranous, connecting 2 parietal placentas, style 1, stigma simple or 2-lobed
Fruit: generally capsule ("silique") with 2 deciduous valves, sometimes breaking transversely or indehiscent
Seeds 1–many per chamber
Genera in family: 300+ genera, 3000+ species: worldwide, especially cool regions; some cultivated for food (especially Brassica, Raphanus ) and ornamental
Recent taxonomic note: Recently treated to include Capparaceae [Rodman et al. 1993 Ann Missouri Bot Gard 80:686–699; Rollins 1993 Cruciferae of Continental North America. Stanford Univ Press]
Family description, key to genera by Robert A. Price.

RORIPPA

YELLOW or WATER CRESS

Annual, biennial, perennial herb; hairs simple, rarely sac-like
Stems 1–many, prostrate to erect, branched or not, often from center of basal rosette, rooting at nodes or not
Leaves: basal, lower cauline sessile or short-petioled, entire to pinnately compound; cauline reduced upward
Inflorescence terminal and lateral
Flower generally small; petals 0 or obovate to narrowly spoon-shaped, pale to bright yellow or white
Fruit linear to round, plump; valves 2 or 4(6); pedicel ascending to recurved, slender, generally with 2 minute glands at base; style 0 or prominent, persistent
Seeds 10–200, (1)2 rows per chamber, dense, generally plump; wing 0; embryonic root at edges of both cotyledons
Species in genus: ± 75 species: worldwide
Etymology: (Old Saxon: for these, perhaps other crucifers)
Reference: [Stuckey 1972 Sida 4:277–340]

Native

R. columbiae (B.L. Rob.) Howell

COLUMBIA YELLOW CRESS

Perennial; hairs spreading, ± dense, short, stiff or longer, pliable
Stem prostrate to erect, generally branched, 1–4 dm
Leaves: basal and lower cauline generally petioled; middle and upper cauline sessile, ± clasping stem, 2.5–5 cm, oblanceolate to oblong, deeply pinnately lobed
Inflorescence axillary elongate
Flower: sepals 2–3.5 mm, ovate to oblong, hairy, persistent in fruit; petals 2.5–4 mm, ± tongue-shaped, yellow
Fruit 3–6 mm, widely oblong; hairs dense; pedicels erect to ascending, often appressed, lower 4–8 mm; style 1–3 mm, hairy, stigma ± expanded
Seed ± < 1 mm, ± reniform, finely papillate
Ecology: Meadows, playas
Elevation: 1200–1800 m.
Bioregional distribution: Modoc Plateau
Distribution outside California: to Washington
Synonyms: R. calycina (Engelm.) Rydb. var. columbiae (B.L. Rob.) Rollins

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