PORTULACACEAE

PURSLANE FAMILY

Annual or perennial herb, generally fleshy
Stems generally glabrous
Leaves simple, alternate or opposite, sometimes stipuled
Inflorescence various
Flower bisexual, radial; sepals generally 2(–8), free or fused at base; petals 3–18, free or ± fused; stamens 1–many, free or inserted on corolla; ovary superior or partly inferior, chamber 1, placenta free-central or basal; styles 2–8, generally fused at base
Fruit: capsule, circumscissile or 2–3-valved
Seeds 1–many, generally black, generally shiny
Genera in family: ± 20 genera, ± 400 species: generally temp Am, Australia, s Africa; some cultivated (Lewisia, Portulaca, Calandrinia )
Reference: [Bogle 1969 J Arnold Arbor 50:566–598]
Family description and key to genera by Dieter H. Wilken & Walter A. Kelley.

LEWISIA

Lauramay T. Dempster

Perennial, generally from short, thick, ± branched taproot, topped by short, sometimes very thick caudex at or below ground level, sometimes from spheric corm
Stem: aerial parts restricted to inflorescence
Leaves generally in basal rosette, simple, entire or not; base wide; margin generally ± translucent
Inflorescence ± scapose; stems 1–many, generally leafless but bracted, sometimes disjointing in age, 1–many-flowered
Flower: sepals 2–8, free, persistent; petals 4–18, variously colored, overlapping in bud; stamens 5–many; styles 2–8, fused at base, stigmas 2–8, thread-like
Fruit: capsule, translucent, spheric or ovoid, circumscissile near base
Seeds 2–many, dark, generally shiny, smooth or finely tuberculate
Species in genus: ± 20 species: w North America
Etymology: (Captain Meriwether Lewis, 1774–1809, of Lewis & Clark Expedition)
Reference: [Elliott 1966 Bull Alpine Gard Soc 34]
Horticultural information: DRN, IRR: pots and rock gardens only; DRY when dormant; DFCLT.

Native

L. cotyledon (S. Watson) B.L. Rob.

CLIFF MAIDS

Root + caudex thick at top, ± slender, branched below
Leaves many, in rosette, 3–9 cm, ovate or spoon-shaped, fleshy, entire or not, tapered to base; tip round
Inflorescence: stems generally 1–6, 10–30 cm, each with a ± flat-topped, 12–many-flowered panicle; flowers exserted from leaves; bracts among flowers and 2–4 below, gland-toothed; pedicels generally < flowers
Flower: sepals 2, ± 1/3 X corolla, round or truncate, margin gland-toothed; petals generally 7–13, ± 15 mm, oblanceolate or obovate, white or pink with darker veins, tip ± notched; stamens 6–9; stigmas 3–4
Chromosomes: n=14
Ecology: Rocky, sandy slopes, subalpine forests
Elevation: 150–2300 m.
Bioregional distribution: Klamath Ranges, High Cascade Range
Distribution outside California: to Oregon, Idaho
Following vars. not clearly distinct, overlap in range. Hybrids with L. leana reported (see L. columbiana).

Native

var. heckneri (C.V. Morton) Munz

HECKNER'S LEWISIA

Leaf: margin dentate
Chromosomes: n=14
Ecology: Rocky cliffs, slopes
Elevation: 300–2100 m.
Bioregional distribution: Klamath Ranges (n Trinity, e Humboldt cos.).