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RUBIACEAE

MADDER FAMILY

Lauramay T. Dempster

Annual, perennial herb, shrub, vine, tree
Leaves generally opposite, entire; stipules generally on stem, sometimes leaf-like (then leaves apparently whorled and stipules considered leaves), adjacent pairs sometimes fused
Inflorescence: cyme, panicle, cluster, or flower solitary, generally terminal and ± axillary
Flower generally bisexual; calyx generally ± 4-lobed, sometimes 0; corolla generally radial, 4-lobed; stamens epipetalous, alternate corolla lobes, generally included; ovary generally inferior, chambers generally 2 or 4, style 1, ± fused if 2
Fruit: 2 or 4 nutlets or a berry, drupe, or capsule
Genera in family: ± 500 genera, 6000 species: worldwide, especially tropical; many cultivated (including Coffea , coffee; Cinchona , quinine; many ornamental)
Reference: [Dempster 1979 Fl CA 4(2):1–47]

GALIUM

BEDSTRAW, CLEAVERS

Annual, perennial herb, sometimes ± shrubby, often ± dioecious, glabrous or hairy, often scabrous
Stem when young 4-angled
Leaves in whorls of 4 or more, including leaf-like stipules
Inflorescence: panicle, or axillary clusters of 1–many flowers
Flower bisexual or unisexual (with sterile stamens or pistils); calyx 0; corolla generally rotate, sometimes ± bell-shaped, generally greenish, fading yellow or white, sometimes reddish, lobes generally 4; ovary 2-lobed, styles 2, ± fused basally
Fruit: 2 nutlets or 1 berry
Species in genus: ± 400 species: worldwide, especially temp
Etymology: (Greek: milk, from use of some species in its curdling)
Hairiness of ovary and fruit generally ± equal on a single plant; staminate plants often identified only by association with pistillate.

Native

G. sparsiflorum Wight

SEQUOIA BEDSTRAW

Perennial, erect, 20–50 cm, loosely tufted, dioecious; base woody
Leaves in whorls of 4, 6–25 mm, broadly elliptic to ovate or round, 3-veined, tip abruptly soft-pointed
Staminate inflorescence: panicle open, leafy; clusters many-flowered
Pistillate inflorescence: flowers generally solitary in axils
Flower: corolla rotate, generally yellowish
Fruit: berry, sometimes hairy
Ecology: Shady places, often in forest
Elevation: 350–2300 m.
Bioregional distribution: Cascade Range Foothills, Sierra Nevada, n Inner South Coast Ranges.

Native

subsp. sparsiflorum

Plant green
Leaf generally elliptic, sometimes ± ovate, thin; puberulent under magnification
Chromosomes: 2n=22
Ecology: Shady places in forest
Elevation: 800–2300 m.
Bioregional distribution: Sierra Nevada.

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