TREATMENT FROM THE JEPSON MANUAL (1993) |
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©Copyright 1993 by the Regents of the University of California
For up-to-date information about California vascular plants, visit the Jepson eFlora. |
AND IS MAINTAINED FOR ARCHIVAL PURPOSES ONLY |
Annual to tree
Leaves simple to pinnately to palmately compound, generally alternate; stipules free to fused, persistent to deciduous
Inflorescence: cyme, raceme, panicle, or flowers solitary
Flower generally bisexual, radial; hypanthium free or fused to ovary, saucer- to funnel-shaped, often with bractlets alternate with sepals; sepals generally 5; petals generally 5, free; stamens (0)5many, pistils (0)1many, simple or compound; ovary superior to inferior, styles 15
Fruit: achene, follicle, drupe, pome, or blackberry- to raspberry-like
Seeds generally 15
Genera in family: 110 genera, ± 3000 species: worldwide, especially temp. Many cultivated for ornamental and fruit, especially Cotoneaster , Fragaria , Malus , Prunus , Pyracantha, Rosa , and Rubus
Reference: [Robertson 1974 J Arnold Arbor 55:303332,344401,611662]
Family description, key to genera by Barbara Ertter and Dieter H. Wilken.
Perennial from short rhizomes and leafless stolons, ± nonglandular
Leaves basal, 1-ternate; leaflet teeth generally simple
Inflorescence: cyme, ± umbel-like, open, 1several-flowered; pedicels recurved in fruit
Flower: hypanthium shallow; bractlets 5; sepals 5; petals 5, ± obovate, generally white; stamens 2035, filaments ± flat; pistils many, ovaries superior, jointed to stout style on side
Fruit: receptacle enlarged, fleshy, red, incompletely covered with achenes
Species in genus: 1530 species: generally n temp
Etymology: (Latin: fragrant)
Reference: [Hancock & Bringhurst 1981 Amer J Bot 68:15]
Hair orientation and plant size have been used to define subspp. but seem to have no taxonomic significance, at least in CA. All species intergrade.
Native |
Often dioecious
Stem generally 520 cm
Leaf leathery; petiole generally 220 cm; central leaflet stalk 110 mm, blade 1060 mm, obovate, densely hairy below, generally glabrous above, rounded to truncate, teeth generally 711, above middle, rounded-obtuse, central tooth << adjacent ones
Inflorescence < or > leaves
Flower generally 2040 mm wide; bractlets unlobed; sepals 610 mm; petals 818 mm
Fruit: receptacle 1020 mm; achene 1.52 mm, smooth
Chromosomes: n=28
Ecology: Ocean beaches, grassland
Elevation: < 200 m.
Bioregional distribution: North Coast, Central Coast
Distribution outside California: to Alaska, also coastal S.America, Hawaii
Cult and possible escape in SCo
Synonyms: subsp. pacifica Staudt
Horticultural information: DRN: 4, 5 &IRR: 7, 15, 16, 17, 22, 23, 24 &SHD: 8, 9, 14, 19, 20, 21; GRCVR; CVS.