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CAMPANULACEAE

BELLFLOWER FAMILY

Nancy Morin, except as specified

Annual to tree
Leaves generally cauline, generally simple, generally alternate, petioled or not; stipules 0
Inflorescence: panicle, raceme, spike, or flowers solitary in axils, generally open; bracts leaf-like or not
Flower: bisexual, radial or bilateral, sometimes inverted (pedicel twisted 180°; hypanthium generally present, ± fused to ovary; sepals generally 5; corolla radial to 2-lipped, generally fused (tube sometimes split down back), lobes generally 5; stamens 5, free or ± fused (anthers and filaments fused into tube or filaments fused above middle); ovary inferior, sometimes half inferior, chambers 1–3, placentas axile or parietal, ovules many, style generally 1, 2–5-branched
Fruit: generally capsule, dehiscing on sides or at tip by pores or short valves
Seeds many
Genera in family: ± 70 genera, ± 2000 species: worldwide. Some cultivated for ornamental (Campanula, Jasione, Lobelia ). Subfamilies sometimes treated as different families.

CAMPANULA

HAREBELL

Annual, perennial herb, from taproot, fibrous roots, or rhizome, glabrous to densely hairy
Stem reclining or erect, branched, 5–150 cm, 4-angled
Leaves cauline, sometimes also basal, generally lanceolate to ovate, thin, fleshy, or leathery, entire to toothed, sessile or petioled
Inflorescence: raceme, panicle, or flowers solitary, terminal or axillary
Flower: corolla cylindric to funnel- or bell-shaped, white to deep blue, lobes linear to triangular; ovary inferior, hemispheric to obconic
Fruit dehiscing by 2–3 lateral pores
Seeds 2 mm, oblong
Species in genus: ± 300 species: n hemisphere; many cultivated, some medicinal
Etymology: (Latin: little bell, from corolla shape)
Reference: [Morin 1980 Madroño 27:149–163]

Native

C. wilkinsiana Greene

WILKINS' HAREBELL

Perennial, forming dense colonies, glabrous; rhizome slender
Stem reclining to erect, 5–30 cm
Leaf 12–20 mm, narrowly oblong, thin; teeth few, sharp; petiole 0
Flower: pedicel 10–30 mm; sepals spreading (erect in fruit); corolla 12–15 mm, funnel-shaped, deep blue, lobes erect; stamens 6 mm, bases ciliate; ovary 4–5 mm, narrowly obconic, style ± 12 mm, blue, upper 50% papillate
Fruit narrowly obconic, weakly ribbed; pores near top
Seed ± 0.7 mm, oblong
Ecology: Wet meadows, streamsides
Elevation: 1800–2600 m.
Bioregional distribution: Klamath Ranges (Trinity Mtns), High Cascade Range (Mount Shasta), n High Sierra Nevada (reported from Deer Creek Meadows, Plumas Co.).

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