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RANUNCULACEAE

BUTTERCUP FAMILY

Dieter H. Wilken, except as specified

Annual, perennial herb, sometimes aquatic
Leaves generally basal and cauline, generally alternate, simple or compound; petioles at base generally flat, sometimes sheathing or stipule-like
Inflorescence: cyme, raceme, panicle, or flowers solitary
Flower generally bisexual, radial; sepals generally 5, free, early deciduous or withering in fruit, generally green; petals 0–many, free; stamens generally 10–many; pistils 1–many, ovary superior, chamber 1, style 1, generally ± persistent in fruit as beak, ovules 1–many
Fruit: achene, follicle, berry, or utricle-like, 1–many-seeded
Genera in family: ± 60 genera, 1700 species: worldwide, especially n temp, tropical mtns; many ornamental (Adonis, Aquilegia, Clematis, Consolida, Delphinium, Erianthis, Helleborus ), some highly TOXIC (Aconitum, Actaea, Delphinium, Ranunculus )
Reference: [Duncan & Keener 1991 Phytologia 70:24–27]

ADONIS

Annual, perennial herb, generally from taproot
Stems 1–few, erect, branched to not
Leaves 2–many-pinnately dissected, cauline; segments generally linear
Inflorescence: raceme or flowers solitary, terminal or axillary
Flower radial; sepals 5, generally ± greenish; petals 5–20, white to yellow or orange; pistils many
Fruit: achene, rough to wrinkled or ridged, beaked
Species in genus: ± 20 species: temp Eurasia; some cultivated as ornamental
Etymology: (Greek: Adonis of mythology, from whose blood the plant allegedly grew)

Introduced

A. aestivalis L.

Annual 3–7 dm, glabrous
Leaves generally 2–3-pinnately dissected, 2–7 cm
Flower: receptacle in fruit 1.5–3 cm; sepals 5–8 mm, ovate to oblong; petals (6)8–10, 8–15 mm, oblong to oblanceolate, yellow to orange, bases bluish to purplish; anthers blue to purplish
Fruit 4–5 mm, 3.5–5 mm wide, with minute, lateral ridge below middle; back keeled; keel base short-toothed; beak 1–1.5 mm, straight
Ecology: Disturbed sites, fields, open pine forest
Elevation: 1200–1400 m.
Bioregional distribution: Modoc Plateau
Distribution outside California: to Washington, Idaho, also e US; native to Europe

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