Jepson Flora Project: Jepson Interchange    

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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California

  • Up-to-date information about California vascular plants is available from the Jepson eFlora.



Alan R. Smith and Thomas Lemieux

Perennial, in soil or on or among rocks; rhizome creeping to erect, scaly
Leaves generally all ± alike (or of 2 kinds, fertile and sterile), generally < 50 cm, often < 25 cm; petiole generally thin, wiry, often dark, in transverse section with vascular strands generally 1–3, less often many in a circle; blade generally pinnate or ± palmate-pinnate (see Adiantum ), often 2 or more compound, lower surface often with glands, ± powdery exudate, hairs, or scales; segments round, oblong, fan-shaped, or otherwise, veins generally free
Sporangia in sori or not, marginal, submarginal, or along veins, sometimes covered by recurved, often modified segment margins (false indusia); true indusia 0; spores spheric, sides sometimes flat, scar with 3 radiating branches
Genera in family: ± 40 genera, 500 species: worldwide, especially dry areas. Definition of Cheilanthes and related genera problematic; traditional limits often untenable.


Plant in soil or rock crevices; rhizome short- to long-creeping-decumbent, generally many-branched, scales generally linear-lanceolate, pale to dark, with darker mid-stripe or not
Leaf < 75 cm; petiole cylindric, reddish brown to blackish; blade generally 2–3-pinnate, generally oblong to narrowly triangular; segments generally small, ± flat or lower side concave (from recurved margins)
Sporangia along margin, in discrete patches to continuous, partly to completely covered by recurved margin (generally not recurved in C. cooperae)
Species in genus: 150+ species: generally Am, generally dry areas
Etymology: (Greek: lip flower, from location of sporangia)
Horticultural information: DFCLT.


C. covillei Maxon

Rhizome short-creeping; scales generally reddish brown, with darker mid-stripe
Leaf 8–22(30+) cm, 2–4(6) cm wide, dark green; petiole < 2 mm wide, scales linear-lanceolate, whitish to reddish brown; blade 3–4-pinnate; segments small, ± round, upper side glabrous, lower concave, obscured by scales that exceed margin, scales originating from axes, > 2 mm, 1 mm wide, ± entire, covering generally more dissected scales
Sporangia obscured by recurved segment margin and scales
Chromosomes: 2n=60
Ecology: Crevices, bases of rocks, sun or shade
Elevation: 600–2400 m.
Bioregional distribution: North Coast Ranges, Sierra Nevada, Sacramento Valley (Sutter Buttes), San Francisco Bay Area, South Coast Ranges, Transverse Ranges, Peninsular Ranges, East of Sierra Nevada, Desert Mountains
Distribution outside California: to Utah, Arizona, Baja California
Hybridizes uncommonly with C. newberryi (C. X fibrillosa L. Underw.) and C. parryi (C. X parishii Davenp.) in s CA; also with C. intertexta.

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