Jepson Flora Project: Jepson Interchange    

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Jepson Interchange (more information)
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Robert F. Thorne

Annual, perennial herb, aquatic, freshwater or marine, bisexual, monoecious, or dioecious
Leaves basal or cauline, alternate, opposite, or whorled, generally ± sheathing at base, glabrous
Inflorescence: cyme; flowers 1–few, subtended by ± sheathing, entire or lobed bract; staminate flowers sometimes deciduous, free-floating
Flower radial; perianth tube 0 or much elongated, peduncle-like in flower; sepals (0)3, green; petals (0)3, colored or white; stamens (1)3–many, generally in 1+ series; ovary inferior, chamber 1, placentas parietal or basal, ovules 1–many; style lobes generally 3, linear, lobed or notched
Fruit: achene or berry-like and dehiscing irregularly, linear to spheric, submersed
Genera in family: ± 17 genera, ± 130 species: worldwide; some cultivated for aquaria, others noxious weeds.


Perennial, rooting at nodes, generally submersed, dioecious or some flowers bisexual
Stem slender, generally branched
Leaves opposite or whorled, generally 3 per whorl, sessile; blade margin minutely, finely toothed
Inflorescence: flowers solitary, axillary, sessile or peduncled; bract generally notched
Staminate flower floating, deciduous or not; perianth tube elongated, slender, peduncle-like; stamens 3–9
Pistillate flower floating; perianth tube elongated, slender, peduncle-like; style slender, stigmas 3, simple or 2-lobed
Fruit cylindric to ovoid
Seeds several
Species in genus: ± 12 species: temp and tropical Am
Etymology: (Greek: of marshes)
[St. John, 1965 Rhodora 67:1–35,155–180]


E. nuttallii (Planch.) H. St. John


Leaves generally not crowded at stem tips; middle and upper 3(4) per whorl; blade generally 6–13 mm, 0.3–1.5(2) mm wide, linear to narrowly lanceolate, tapered to acute tip
Staminate inflorescence: bract ovoid, abruptly pointed; flower deciduous, free-floating
Staminate flower: sepals ± 2 mm, ± 1.5 mm wide; petals generally 0 or ± 0.5 mm
Pistillate flower: sepals ± 1 mm
Seed short-soft-hairy
Chromosomes: 2n=48
Ecology: Shallow water, streams, lakes, ponds, ditches
Elevation: 500–2800 m.
Bioregional distribution: Klamath Ranges, High Sierra Nevada, Sacramento Valley, San Bernardino Mountains, Great Basin Floristic Province
Distribution outside California: to Washington, c US
Flowering time: Jul–Aug

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